DDSH Thuc Vat Rung Thong 3 La Lam Dong

B GIO DC V O TO TRNG I HC LT
BO CO TI KHOA HC CP B
NGHIN CU A DNG SINH HC THC VT RNG THNG BA L (PINUS KESIYA) MC T NHIN LM NG V VNG LN CN
M TI B 2009 14 30 CH NHIM TI: ThS. NGUYN DUY CHNH
NHNG NGI THC HIN: -TRN VN TIN, Trung tm Nghin cu Lm Sinh Lm ng -HUNH TH NGC NH, Trng i hc Ph Yn -VNG THC LAN, Trng i hc Lt -NGUYN VN NGC, Trng i hc Lt -HUNH TH BNH, Trng i hc Lt
TM TT S dng phng php xp chng xc nh din tch thch hp ca tiu chun nghin cu a dng sinh hc kiu rng thng ba l mc t nhin Lm ng v vng ln cn.Cc xp chng c kch thc: 10m x 10m, 15m x 15m, 20m x 20m, 25m x 25m, 30m x 30m, 35mx 35m, 40m x 40m. tiu chun c xc nh vi knh thc 35m x 35m l thch hp v thc t nht cho cc nghin cu a dng sinh hc thc vt, c bit l ng vi rng thng ba l (Pinus kesiya) mc t nhin Lm ng v cc vng ln cn trn cc ai cao t 800m n 2000m. Trong cc nghin cu ca chng ti, tng s tiu chun c thc hin l 20 , mi tiu chun u c ghi nhn ta a l (v , kinh ), cao, dc, hng dc, thnh phn loi, dng sng, s lng c th v ch s a dng Margalef. T cc kt qu chng ta c c ch s c trng chung v a dng sinh hc (ch s trung bnh), thnh phn loi v a dng dng sng ca kiu rng ny. Ch s Margalef trung bnh (DMarg) c ch ra l 3,76. Thnh phn loi kh giu v a dng, bao gm 244 loi thuc 179 chi, 68 h ca 4 ngnh thc vt mch (Lycopodiphyta, Poplypodiophyta, Pinophyta. Magnoliophya). C 8 loi c ghi trong sch Vit Nam. C 8 dng sng trong : Megaphanerophytes (0,82) Microphanerophytes (9,01) Nanophanerophytes (18,44), Chamaephytes (27,46), Therophytes (27,05), Lianophanerophytes (6,15), Cryptophytes (6,65), Epiphytes (4,51). T kha: xp chng, tiu chun, a dng sinh hc, thng ba l, ch s Margalef, thnh phn loi, dng sng, Lm ng.
SUMMANY The paper present a method of using superposed plots to identify the most suitable area of perquadrates for the research into the biodiversity of the natural three leaved Pine forests distributed among Lam Dong and its subareas. The sizes of the superposed plots are: 10m x 10m, 15m x 15m, 20m x 20m, 25m x 25m, 30m x 30m, 35m x 35m, 40m x 40m. The 35m x 35m sized perquadrates are suggeted to be the most appropriate and most pratical for researchs of plant biodiversity, especially thoes of the three leaved pine forests (Pinus kesiya) naturally grown in Lam Dong and the its subareas with the altitude from 800m to 2000m. In our researches, the total number of perquadrates is 20. Each perquadrate has a record of its own indices including: geological co-ordinate (latitude and longitude), alttitude, gradient, klinogeotropicsm, species composition, life form, number of individuals, Margalef index. From the results we have specific index of biodiversity for natural three-leaved pine forests, species composition and diversity of the life form in the forests. The everage Margalef index (DMarg) is 3,76.The species composition is rather rich and diverse, including 244 species, 179 genera,68 families of 4 vascular plant phyta (Lycopodiophyta, Polypodiophyta, Pinophyta, Magnoliophyta). There are 8 species which have been recorded in Red Data Book of Viet Nam. There are 8 life forms, including: Megaphanerophytes (0,82%), Microphanerophytes (9,01%), Nanophanerophytes (18,44%), Chamaephytes (27,46%), Therophytes (27,05%), Lianophanerophytes (6,15%). Cryptophytes (6,56), Epiphytes (4,51%). KEY WORDS: Superposed plot, Perquadrate, Biodiversity, Margalef Index, Three leaved pine forest, Species composition, Life form, Lam Dong.
M U Ngy nay a dng sinh hc tr thnh mi quan tm ln ca con ngi trn khp hnh tinh. Vit Nam c nhiu cng trnh nghin cu v a dng sinh hc cc cp khc nhau. Nhng nghin cu v a dng di truyn ca Nguyn Hong Ngha, Nhng nghin cu v a dng loi ca Nguyn Tin Bn, Nguyn Ngha Thn, Nhng nghin cu v a dng h sinh thi ca Thi Vn Trng v c nhng cng trnh s ca Phm Hong H. Tt c chng t Vit Nam l nc a dng v sinh vt. S giu c v sinh vt c tng cha trong cc kiu thm thc vt khc nhau. Lm ng l tnh min ni vi nhiu kiu a hnh cc ai cao khc nhau, c cc kiu thm thc vt khc nhau trong kiu rng tha cy l kim ca loi thng ba l (Pinus kesiya) l in hnh nht cho thm thc vt ni y. Rng thng ba l Lm ng c khong 192,320 ha, trong khong 148.000 ha l rng thng mc t nhin cc ai cao t 800 m n 2.000m. S cng trnh nghin cu c lin quan n a dnh sinh hc rng thng y cn t v nhn chung cha phn nh a dng thc vt ca kiu rng ny. c c kt qu v a dng thc vt ca rng thng, cn thit lp cc tiu chun. Song kch thc ln va ca l bao nhiu th thch hp, th phn nh tnh a dng sinh hc ca kiu rng ny. l vn cn t ra khi ta phi nghin cu cc a hnh chia ct mnh, c dc ln. ng thi cn thit lp nhiu trn nhiu cao kt qu nghin cu phn nh khi qut nht a dng sinh hc thc vt ca kiu rng thng ba l Lm ng.
PHN 1: TNG QUAN TI LIU I. S LC V IU KIN T NHIN CA LM NG 1. V tr a l. Lm ng l tnh min ni nm pha Nam Ty Nguyn, l u ngun ca 4 h thng sng ln: Sng ng Nai (ng Nai) S r pok (c Lk) Sng Ly (Bnh Thun) Sng Ci (Ninh Thun) Lm ng c v tr a l nh sau: Pha Bc gip tnh k Lk Pha Ty Bc gip tnh k Nng Pha Ty v Ty Nam gip tnh ng Nai v tnh Bnh Phc Pha Nam gip tnh Bnh Thun Pha ng gip tnh Ninh Thun v Khnh Ha Lm ng nm gia cc ta a l: T 11012/47// n 12019/01// v Bc T 107016/23// n 108042/11// kinh ng Theo nin gim thng k nm 2005 ca tnh Lm ng th din tch ton tnh l 977.219 ha, chim khong 2,9% din tch c nc. 2. a hnh a th. a hnh l mt hnh th phn nh yu t a cht v qu trnh a mo, do gn lin vi ngun gc a cht v tui khu vc, a hnh tnh Lm ng nhn chung thuc dng vng ni, t ni thp, ni trung bnh n ni cao. cao ni thay i t 200m n 2.200m. a hnh Lm ng nghing dn t hng ng Bc xung hng Ty Nam. Nh vy c im ni bt ca a hnh Lm ng l nghing, to nn s phn bc r rng, to nn cc ai i cao khc nhau vi rt nhiu nh ni cao nh Bidoup (2.287m), Langbiang (2.167m), Ch You Kao (2.006m), MNeun Ro (1996m), Be Nom Dan Seng (1.931m), Braion (1.874m), Quan Du (1.805m), Ch Yn Du (1.784m), M Neun Pautar (1.664m), MNeun Lamleo (1.623m). Trong mi quan h vi a cht a mo, c th phn chia a hnh ca tnh Lm ng ra cc dng sau:
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+ a hnh thung lng: Gm cc b mt tng i bng phng, t dc, c ngun gc tch t thung lng gia ni hoc cc bi tch sng sui hin i. t y ty thuc vo ngun gc mu th v mc bo ha nc m c xp vo cc loi t ph sa, dc t hoc t gley v hu ht c kh nng thch hp cao cho s pht trin ca nhiu loi cy dng ha tho, cy mt nm hay cy ti sinh chi lu nm. + a hnh i ni thp n trung bnh: y l kiu a hnh c cc di i ni t dc, thng th dc c dc nh di 200 v c cao nh di 800m n 1.000m. dng a hnh ny phn nhiu l cc i ni c ngun gc phun tro bazan vi nn t nu hoc nu vng trn m bazan. + a hnh ni cao: y l kiu a hnh cc khu vc ni c cao trn 800m, thng c dc mnh trn 200, l kiu a hnh chia ct mnh. Ch yu l cc khu vc c ngun gc xm nhp JuraCreta (Granit, Dacit hoc Andezit), hay cc trm tch Mesozoi (phin sa, phin st). dng a hnh ny ph bin l cc loi t vng , vng hay t xm trn cc magma acit trung tnh hoc phin. t y phn ln l c tng mng. Chnh do c dc ln, c nhiu vng dc trn 300, li c tng t mng, cho nn a hnh ni cao ch thch hp cho cy trng lm nghip, cy g v phn ln din tch y c rng che ph. 3. Kh hu thy vn. Tnh Lm ng c ch kh hu nhit i gi ma. Tuy nhin nh phn trn cp, tnh Lm ng c din tch ln, c chia thnh nhiu kiu a hnh vi cc ai cao rt chnh lch nhau t 100 n 2.200m. y c 2 Cao Nguyn ln l Cao nguyn Langbiang v Cao nguyn Di Linh Bo Lc. Cao nguyn Langbiang hay cn gi l Bnh Sn Langbiang c bao bc bi cc dy ni hnh cnh cung v pha Ty, Bc v ng c cao khong 2.000m. Cao nguyn ny c dng thung lng v thp dn v pha Nam. Cn Cao nguyn Di Linh Bo Lc c dng thung lng c tri trn din tch rng ln vi cao trung bnh t 800 n 1.000m, xen ln l mt s dy ni cao nh Braian. Chnh do a hnh tng ai cao vn b chia ct rt mnh bi cc khi ni ln, nh, Lm ng li c nhiu ai cao, vi chnh lch cao ln, cho nn tuy nm trong vng kh hu nhit i gi ma nhng ch kh hu bao gm ch bc x, ch nhit , ch m v lng ma, s ngy ma, lng bc hi, s gi nng l khng ng nht. a) Ch bc x Bc x tng cng Bo Lc l 128,0 kcal/cm2/nm. Trong khi bc x tng cng Lin Khng l 139,9 kcal/cm2/nm.
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b) Nhit khng kh Nhit khng kh trung bnh nm Lt l 17,90C. Trong khi nhit trung bnh nm Bo Lc l 21,90C, Lin Khng l 21,20C. Nhit ti cao Bo Lc l 33,50C, cn Lt l 29,40C. Nhit ti thp Lt l 4,90C, cn Bo Lc l 5,50C. Nhn chung nhit chnh lch nhau gia cc thng tng vng l khng nhiu, nhng chnh lch nhit ngy m li cao, c bit l Cao nguyn Langbiang, ch nhit cng thun li cho s pht trin ca nhiu loi thc vt, c bit ch nhit 2 Cao nguyn: Cao nguyn Di Linh Bo Lc v Cao nguyn Langbiang rt ph hp cho s pht trin cc loi cy l kim, trong c loi thng ba l (Pinus Kesiya). c) Ch ma Lng ma trung bnh nm a Th l 3.790mm, Bo Lc l 2.722mm, Di Linh l 1.721mm, Lt l 1 1.821mm, Lc Dng l 1.811mm. S ngy ma trung bnh Bo Lc l 191 ngy, Lin Khng l 160 ngy, cn Lt l 167 ngy. Nhn chung ch ma cho php Lm ng pht trin nhiu kiu thm thc vt ca vng ma nhit i. S ngy ma cc thng 7, 8, 9, dao ng t 25 - 28 ngy. Cc thng 1, 2 s ngy ma vt khng qu 4 ngy, cc thng cn li s ngy ma trung bnh vo khong 4 n 20 ngy. Ma ma tp trung nhiu t thng 5 n thng 10. Ma kh t thng 11 n thng 4. Trong ma kh, do nh hng ca gi ma ng Bc nn Lm ng ma rt t, lng ma ch chim 10 15% lng ma c nm. Chnh c im ny to iu kin cho s pht trin kiu rng tha cy l kim c loi thng ba l (Pinus Kesiya) l u th. d) m khng kh Do c im v v tr a l v a hnh m m cc vng khc nhau trong tnh Lm ng khc nhau. m tng i ca khng kh vo ma ma kh cao (80 90%), vo cc thng 6, thng 7, thng 8 v thng 9 m khng kh l ln nht (90%). Cc thng ma kh m khng kh khong 65 75% Lt, 70 78% Lin Khng. e) nh sng nh sng l mt trong cc yu t sinh thi nh hng rt ln n cc hot ng sng, quy nh phn b ca cc loi thc vt. Ch chiu sng cng khc nhau cc vng khc nhau Lm ng.
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S gi nng trong nm tnh trung bnh cho vng Bo Lc l 1.988 gi. Trong khi Lt l 1.868 gi, cn Lin Khng l 2.268 gi. S ngy c sng m Bo Lc l 85 ngy, Lin Khng l 30 ngy. Lm ng l ni c ngy ngn, vo cc thng khc nhau cng c thi gian chiu sng l khc nhau. T thng 4 n thng 9 thi gian chiu sng trung bnh trong ngy l 9 10 gi, cc thng khc thi gian chiu sng l 10 12 gi. Nhn chung ch chiu sng ph hp cho vic pht trin thm rng thng ba l l loi cy a sng. f) Sng sui v thy vn Nm trong khu vc a hnh ni cao chia ct mnh v c lng ma ln, nn mng li sng sui Lm ng kh phong ph. S lng sng sui c chiu di vt trn 10km trong phm vi ton tnh ln ti gn 60km. Trong c mt s sng, sui ln nh sng ng Nai, sng a Nhim, sng Dng, sng Tam, sng i Nga, sng Th, sng Tamat, sui Katch, sui anian, sui a Nhau, sui a Plaite Mt li sng sui thay i khong 0,28 1,1km chiu di/km2 vi din tch theo thng k khong 15.500ha. Sng sui Lm ng nhn chung c bc thm sng hp, sn dc, nhiu thc ghnh, dng chy mnh v phn b khng u trong nm. Modul dng chy ton nm dao ng t 18 20/s/km2, vo ma ma l thng t thng 7 n thng 11, lu lng mt s sng sui chnh ng vi tn sut 1% ln n 1.000 5.000m3/s, gy l lt nghim trng. Trong khi vo ma kit, t thng 1 n thng 3, modul dng chy kit ch t 0,25 n 9,1/s/km2, hn ch n kh nng cung cp nc cho thc vt. Lng dng chy trung bnh mi nm mi khu vc ty thuc vo lu vc, lng ma, a hnh v a cht, c s khc bit r rt. Vng Bo Lc Hoai 39 40/s/ km2 Vng Lt c Trng 23 28/s/ km2 Vng n Dng 23 24/s/ km2 Lng dng chy kit do cn ph thuc vo mc che ph ca thm thc vt. Lng dng chy s nh hng n tnh hnh xi mn. Lm ng l mt tnh min ni, c a hnh kh phc tp, a hnh nhiu ni chia ct mnh, dc ln, v vy t d b xi mn, ra tri, nht l cc vng khng c rng che ph. Hin tng xi mn ph bin l xi mn khe, rnh trn din tch hp.
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L lt thng xy ra vng h lu sng ng Nai gy ng lu di cho cc vng Ct Tin, Hoai, Th, ni c a hnh rt thp trng. 4. Th nhng Tnh Lm ng c din tch 977.219 ha vi 8 nhm t nh sau: + Nhm t ph sa (Flavisols) y l nhm t c to nn do s bi lng ca sng sui. Tnh cht t thay i ph thuc vo sn phm phong ha ca mu cht to thnh t vng thng ngun ca tng lu vc, thi gian v iu kin bi lng. Nhm t ny gm: t ph sa chua, t ph sa giu mn, t ph sa gly. + Nhm t gly (Gleysols) Nhm t ny c hnh thnh vng a hnh trng, mc nc ngm nng, to iu kin thun li cho qu trnh kh t. t co mu xanh, c ngun gc thy thnh. Nhm ny gm t gly ng nc nhn tc, t gly c tng si sn nng, t gly giu mn, t gly c tng mt giu mn. + Nhm t mi bin i (Cambisols) Nhm t ny c hnh thnh trong iu kin ra tri, feralit ha, gley ha ang mc thp ca cc qu trnh . Nhm t ny gm: t mi bin i ng nc t nhin, t mi bin i chung t mi bin i giu mn, t mi bin i tng mng ng nc. + Nhm t en (Lavisols) Nhm t ny c hnh thnh do qu trnh ra tri tch ly st. Nhm t ny gm: t en giu mn, t en chua, t en gly c tng vng loan l. + Nhm t Bazalt (Ferralsols) y l nhm t chim din tch ln Lm ng. Nhm t ny c hnh thnh do phong ha khong st, hnh thnh cc khong hot tnh thp, khng c kh nng phong ha tip nh Kaolinit, tch ly cc oxit Fe, Al Nhm t ny gm: t chua giu mn, t chua ngho baz, t chua tng mt giu mn, t si sn su, t si sn nng.
+ Nhm t xm (Acrsols) y l nhm t ln nht, chim n 2/3 din tch t ton tnh Lm ng, phn b hu ht cc huyn trong tnh, t a hnh ni cao n a hnh g i thp trng v thung lng, trn cc loi m. Nhm t ny gm nhiu loi t nh: t xm vng, t xm c tng thm mc, t xm giu mn tch Al, t xm gly, t xm ngho baz, t xm rt chua vng, t xm tng mt giu mn, t xm tng mng. + Nhm t mn trn ni cao (Alisols) Nhm t ny gp trn ni cao trn 2.000m v vy din tch nh. Vi c trng c hnh thnh trong iu kin nhit thp, m cao quanh nm. + Nhm t b xi mn mnh (Leptosols) Nhm t ny c trng l dy tng t rt mng, t mn, ch yu vng g i, din tch rt nh. 5. t rng v rng Tng din tch t nhin tnh Lm ng t 1.017.260 ha vo nm 1992, nay iu chnh cn 977.219 ha (Theo nin gim thng k nm 2005 ca tnh Lm ng). Nh vy din tch gim 40.041 ha, nguyn nhn l do thc hin ch th 364/CP ngy 06/01/1991 ca Chnh ph v Cng vn s 341/TCCP P ngy 03/10/1996 ca Ban t chc chnh ph v vic cng nhn h s bn a gii hnh chnh cc cp tnh Lm ng, theo iu chnh mt s din tch t sang cho tnh k Lk, tnh Ninh Thun, tnh Bnh Thun. Din tch t c rng ca tnh Lm ng l 607.280 ha, trong din tch rng t nhin l 557.857ha v din tch rng trng l 49.423 ha. Nh trn cp, do iu kin t nhin, c im v a hnh phong ph, ch kh hu, thy vn a hnh th cho php Lm ng pht trin nhiu kiu thm thc vt khc nhau vi nhiu kiu rng khc nhau: Rng kn thng xanh ma ma nhit i. Theo Richard th nhng vng lnh th nm vnh ai nhit i thng c lng ma cao, v khi lng ma vt qu gii hn 1.500mm trong nm, s cho php cc thc vt pht trin mnh v to ra kiu rng ma nhit i. phn trn cp, tt c cc vng Lm ng u c lng ma ln, vt xa gii hn ny, thng th t 1.800mm n 2.400mm nm. V li u thuc ai v 110 v Bc. V vy kiu rng kn thng xanh ma ma nhit i gp ph bin Lm ng, tt cc ai cao, tt c cc a phng (cc huyn) u c s che ph ca kiu rng ny.
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Rng kn thng xanh ma ma nhit i vi cc c trng rt a dng v thnh phn loi, rt a dng v dng sng, cu trc phn tng gm nhiu tng. V thnh phn loi trong mt qun x c n vi trm loi thc vt, thuc v vi chc h ca nhiu ngnh thc vt nh: Bryophyta, Lycopodiophyta, Equisetophyta, Polypodiophyta, Gymnospermae, Angiospermae. Cc loi cy c th ch gp cc vng thp hay ni va nh cc loi thuc h du (Dipterocarpaceae), h Bng Lng (Lythraceae). Hay nhiu loi ch gp phn b cc ai cao, nh cc loi h Quyn (Ericaceae), h Ch (Theaceae) hay cc loi h D (Fagaceae), cc loi h Long No (Lauraceae). V cu trc qun x thc vt ca kiu rng ny thng c cu trc 5 tng: - Tng g ln (A1): Nh Trm (Syzigium), Bi li (Litsea), D (Lithocarpus, Quercus, Castanopsis), Cm (Elaeocarpus) - Tng g nh (A2): G Tm (Ostoides), M nng (Marcaranga), Th rng (Diospyros), Ficus, Fagaceae, - Tng g nh (A3): Mt s loi thuc Rubiaceae, mt s loi thuc Myrtaceae, Melastomataceae, Ericaceae, Theaceae, - Tng cy bi (B): Gm nhiu loi thc vt ca c lp Magnoliopsida lp Liliopsida nh: Cau chut (Pinanga), My (Calamus), mt s loi cy h cy C ph (Rubiaceae), h Qu n (Acanthaceae), h Nhn sm (Araliaceae), loi u rng (Thevesia Palmata) - Tng c (C): Vi c trng l tha tht, tng ny khng ph kn b mt, che ph khong 10%. Tuy nhin cc loi cy cng kh a dng nh: Quyn b Selaginella thuc ngnh Lycopodiophyta, Ru tc trng Leucobrium thuc ngnh Bryophyta, cc Dng x chi Pteris, Adiantum, Antrophyum, Crypsinus thuc ngnh Polypodiophyta, nhiu loi thuc magnoliophyta. Bn cnh , an xen vi cc loi cy v cc tng, c tng dy leo thuc mt s h thc vt nh Cucurbitaceae, Vitaceae kiu rng thng xanh ma ma nhit i ny cn c c trng bi cc hin tng : hin tng cy c bnh v nh cy Sloan (slonea) , cy Cm (Elaeocarpur) thuc h Elaeocarpaceae, mt s loi h o ln ht Anacardiaceae hin tng bp c cng rt ph bin kin rng ny. Rng kn thng xanh hn giao cy l rng, l kim. Kiu rng kn thng xanh hn giao cy l rng, l kim ma ma nhit i pht trin trn ni cao thng gp trong cc rng trn ni cao, c ai cao cao trn 1000m, cng ln
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cc ai ni cao kiu ny cng thng gp. c bit l cc vng ni cao trn 1.600m, m in hnh l vn quc gia Bidoup - Ni B. vn quc gia Bidoup - Ni B u tin phi k n kiu rng hn giao gia cc loi cy ht trn l thng hai l gip (ucampopinur krempfii) thuc h Pinaceae, loi thng nm l (Pinus dalatensis) ri k n loi Pmu (Fokienia hodginsii), loi Hng Tng (acridium Pierei) v mt s loi cy l kim khc hn giao vi cy l rng. y l nhng kiu rng nguyn sinh rt c bit, rt c ngha v mi trng, c tri trn a hnh ni v ni cao rng ln, l u ngun ca nhiu dng chy, nhiu con sng quan trng. Rng kn thng xanh vi rng hn giao cy g v rng tre na. y l kiu rng th sinh do cc loi tren na xm ln rng g, thng th kin rng ny phn b nhiu vng thp, ni m, ven sui. kiu rng ny cy g c th mc ri rc to thnh tng ring, tng kia l tre na. Kiu rng ny gp nhiu Hoai, Th, Ct Tin, Lm H, Di Linh Rng tre na thun loi Rng tre na thn loi phn b ni m, ven sui, ch yu gp cc vng c ai cao di 1000m, tp trung Bo Lc, Di Linh, Haoai, Th, Ct Tin. Rng tha cy l kim a sng cha hn in hnh ca kiu rng cy l kim a sang chu hn Lm ng l kiu rng thng ba l (Pinus kesiya). Rng thng ba l phn b rng trn cc kiu a hnh khc nhau, cc ai cao khc nhau t 800m n 2.000m. Kiu rng thng ba l to nn nt c trng cho kiu thm thc vt Lm ng, c th ni Lm ng l th ca loi thng ba l Vit Nam. So vi kiu rng kn thng xanh th kiu rng thng ba l c thnh phn loi khng phong ph bng, kiu rng thng ba l ch c trn 200 loi, thuc vi chc h thc vt. V cu trc phn tng kiu rng ny cng n gin hn kiu rng kn thng xanh cy l rng, rng thng ba l ch c 3 tng. Tng g ln ch c loi thng 3 l l u th. Tng g nh c vi chc loi thuc mt s h thc vt nh: Cp mc hnh sao (Craibiodendrom stellatum), Nem l liu (Vaccinium iteophyllum), Quyn hoa trng (Pierris ovalifolia) thuc h Ericaceae. Loi thn tn (Aporusa serrata), Sc Dalton (Glochidion daltonii), Me rng (Phyllanthus embrica) thuc h Euphorbiacea. Loi Quercus serrata, Quercus lanata, Quercus setulosa, Lithocarpus dealbathus thuc h Fagaceae. Loi qun hoa (Helicia) h Proteaceae. Loi du ru (Myrica esculenta) h Myricaceae
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Tng c bi c trn 200 loi ca hng lot h thc ca c lp Liliopsida, Magnoliopsida ca ngnh Magnoliophyta, v ca cc ngnh Lycopodiophyta, Polypodiophyta Pinophyta. Nh cc loi Mua lng (Melastoma saigonensis), cc loi ca h Asteraceae, Fabaceae, Poaceae Bn cnh kiu rng tha cy l kim ca loi thng ba l (Pinus kesiya) chim din tch ln, phn b rng ri Lm ng, cn gp kiu rng thng hai l (Pinus merkusii) vi din tch khng ln. Thng hai l cn c th hn giao vi loi du Tr ben (Dipterocarpus obtusifolius) thuc h Dipterocarpaceae to nn kiu rng tha hn hp giao cy l rng l kim. Rng trng Kiu rng Lm ng c th trng nhiu mt s loi cy g nh: Bch n, Trm bng vng, keo tai tng, du. tuy nhin phn ln din tch rng trng Lm ng l rng trng loi thng ba l. II. TNH HNH NGHIN CU A DNG SINH HC V A DNG THC VT RNG THNG BA L VIT NAM V LM NG. 1. Tnh hnh nghin cu a dng sinh hc Vit Nam Vit Nam c coi l l nc c a dng sinh hc cao, l mt trong 10 trung tm a dng sinh hc quan trng ca th gii v c th hin qua s phong ph ca ngun gen, s lng loi, cc kiu cnh quan, h sinh thi v vng a l sinh hc. T ngn xa cho n nay, a dng sinh hc ca Vit Nam v ang gp phn quan trng khng ch i vi s tn ti v pht trin ca th gii sinh vt m c vi s pht trin kinh t, x hi, vn ha, gio dc ca con ngi nh: Cung cp lng thc, thc phm, thuc cha bnh, ngun nguyn liu cho nhiu ngnh cng nghip. Bn cnh gi tr kinh t, a dng sinh hc cn c gi tr sinh thi, mi trng ln nh: iu ha ngun nc, iu ha kh hu, chng nhim mi trng Mt gi tr quan trng khc ca a dng sinh hc l gip cho con ngi to nn nhng nt p v o c, gio dc thm m, ngh ngi, dng sc v tn ngng ca mnh. Tri qua nm thng, cng vi nhng bin c ca lch s v chnh tr, kinh t v x hi, a dng sinh hc ca Vit Nam v ang b suy thoi v s suy thoi ny ang din ra vi tc rt nhanh trong nhng nm gn y. C nhiu nguyn nhn dn n s suy thoi a dng sinh hc ca Vit Nam, song nguyn nhn c bn nht l mt rng t nhin. Khai thc qu mc v nhn thc v vai tr, gi tr ca a dng sinh hc cn hn ch. Cui cng, ci ngun ca mi nguyn nhn suy thoi a dng sinh hc l do mu thun gia kh nng cung cp ti nguyn sinh vt v nhu cu s dng ca con ngi.
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Nh vy a dng sinh hc l mt trong nhng ti nguyn qu gi m khng th ly cc gi tr khc thay th c, a dng sinh hc tr thnh c s cho s tn ti v pht trin ca con ngi v loi ngi. Nhn thc c gi tr ti nguyn thin nhin, thin nhin ca a dng sinh hc, loi ngi cng nhn thc n s cn thit phi bo tn a dng sinh hc, tng cng v pht trin ngun ti nguyn a dng sinh hc nhm p ng nhu cu ngy cng gia tng ca con ngi trong thi hin i. lm c chuyn khoa hc v a dng sinh hc cng ngy cng pht trin. Vy th cm t a dng sinh hc, hay ni cch khc hn l thut ng a dng sinh hc c khi nguyn t u. Nu nh trong phn loi hc, loi l n v c s (c bn) ca phn loi, th trong nghin cu a dng sinh hc, loi cng c hiu nh n v c s l gc gc cho mi hot ng nghin cu a dng sinh hc. Cc nh phn loi hc t cc th k trc, khp cc ni trn th gii tin hnh cc hot ng phn loi, to nn cc h thng phn loi, phn chia v cng b cc loi trong cc h thc vt khp cc nc trn th gii. n nay loi ngi c mt h thng tri thc khng l v cc h sinh vt mi ni trn th gii v ring Vit Nam cng vy. V chnh cc h thng tri thc v cc h thc vt, h ng vt, h sinh thi l tri thc v a dng sinh hc. Ni nh vy khoa hc v a dng sinh hc c mm mng v ly khoa hc phn loi lm nn mng. Vit Nam, ngay t nhng nm 1879 1899, Pierre vit cun Thc vt ch rng Nam B vi dung lng 400 trang c xut bn Pari. K hng lot cc tc gi ngi Php nh Petelot, Poilane, Chevalier, Gagnepain dy cng nghin cu h thc vt Vit Nam t nhng nm 1907 n 1937 v cho ra i b Thc vt ch i cng ng Dng gm 7 tp, c xut bn Pari do H.Lecomte ch bin. Trong b Thc vt ch i cng ng Dng c rt nhiu loi thc vt bc cao thuc hng lot cc h thc vt c m t k lng. B sch ra i cng cc bo tng thc vt lu tr cc mu vt thu thp Lm ng tr thnh nn tng cho nhng cng cuc nghin cu thc vt Vit Nam cho cc tc gi v sau , ch yu l cc tc gi ngi Vit Nam. Cc bo tng hin vn lu tr cc mu vt , tr thnh kho t liu qu gi nh: Bo tng thc vt thnh ph H Ch Minh 85 Trn Quc Ton, Phng Bch tho thc vt thuc i hc Tng hp H Ni (nay l trng i hc Bch khoa t nhin thuc i hc Quc gia H Ni). Sau ny, t nm 1960 n 2001, hng lot cc tc gi khc tip tc nghin cu v b sung cc h cha c Thc vt ch i cng ng Dng cp. Cc tc gi cho ra cc tp khc nhau trong b H thc vt Cambt, Lo v Vit Nam do A.Aubreville
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v J.Leroy, Ph.Morat l ch bin, trong cp, b sung hng lot cc loi, cc h thc vt cho h thc vt Vit Nam. Nhng nm t 1969 n 1976 di s ch bin ca L Kh K, nhiu nh nghin cu thc vt nh V Vn Chi, Phan Nguyn Hng, L Kh K cng b 7 tp b Cy c thng thy Vit Nam. Trong khi t nm 1970 n 1972 Gio s Phm Hong H cng b b Cy c min Nam Vit Nam gm 2 tp. Sau ny Gio s Phm Hong H li cng b tip 3 tp ca b Cy c Vit Nam vo nm 1991 1993 ti Montreal. n nm 1999 2000 nh xut bn tr thnh ph H Ch Minh ti bn b Cy c Vit Nam ca Gio s Phm Hong H. y l b sch s, c nhiu ngi nghin cu thc vt c cc lnh vc lin quan n thc vt tra cu mt cch rng ri v thng xuyn trong b Cy c Vit Nam; hu ht cc h thc vt bc cao ca cc ngnh: Psilophyta, Lycopodiophyta, Equisetophyta, Polypodiophyta, Pinophyta, Magnoliophyta c tc gi cp. Trong tng loi c m t c km theo hnh v v nhiu thng tin lin quan n loi m tc gi dy cng tp hp qua qu trnh lu di trong cc nghin cu ca ng, trong cc thu thp tri thc bn a, trong cc thu thp tri thc ca nhn loi. Theo Gio s Phm Hong H, Vit Nam l nc c h thc vt phong ph vo bc hng u ca th gii vi khong 12.000 loi thc vt. Trong s cc loi thc vt , rt nhiu loi c ch ra c phn b Lm ng, hay phn b a danh no thuc Lm ng, nh phn b Lt, phn b Bo Lc, Di linh, n Dng hay phn b Langbiang Cng tromg s c loi tc gi ch ra phn b trong kiu rng thng. V d: Craibiodendron henryi w.w.sm.var.bidoupensis Smitin.&Pham Hoang Phn b: Lm ng 2.000m Craibiodendron stellatum (Pierre) w.w.sm. l loi cp mc hnh sao Phn b: Vng ni cao, Lt, Langbiang. Hay loi chn voi mm (Elephantopus mollis H.B.K.) Phn b: Rng tha, rng thng, da l, thng thng I XII. L non c n cha cht nn ung thu, bch huyt, c ch vi khun, tr bnh phi. ng thi tc gi cng ch ra tn ng ngha ca loi, v d: Elephantopus mollis H.B.K. (E.bodinieri gagn. E.tomentosus Kosternon L..) Bn cnh , trong qu trnh nghin cu h thc vt rng thng ba l (Pinus kesiya) tuy l mt h thng tht vt phc tp, khng tht qu giu loi v vi cu trc rng kh n gin, song cng cn rt nhiu tra cu t liu, k c nhng t liu khng vit khi qut cho h thc vt Vit Nam m mang tnh chuyn nh: B Cy g rng Vit Nam ca Vin iu
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tra quy hoch rng gm 7 tp, hay cun sch Cy thuc v v thuc Vit Nam ca Tt Li, hay cun Dc in cy thuc Vit Nam ca V Vn Chi, l nhng ti liu cng rt hu ch cho tra cu, cho so snh khi gp nhng Taxon kh nh loi. Li c nhng ngun ti liu rt gn vi nhng nghin cu ca chng ti nh Danh lc thc vt Ty Nguyn do Nguyn Tin Bn ch bin, vi s tham gia ca nhiu tc gi cho cc ngnh, cc h nh Phan K Lc, Trn nh L, Nguyn Tin Hip, Nguyn Th , L Kim Bin Cc loi thuc cc h thc vt khc nhau c dn ra di tn gi khoa hc, tn a phng v ni phn b c ch r nhiu a phng Ty Nguyn v Lm ng, ch r n cc a danh c th cc x hay mt ngn ni no m cc tc gi dy cng nghin cu, thu thp mu vt trong mt s nm. hon thin chnh xc v danh php ca cc loi thc vt khng th khng nhc n ti liu tra cu di nhan Danh lc cc loi thc vt Vit Nam gm 3 tp. Tp 1 do Trung tm nghin cu ti nguyn v mi trng, i hc Quc gia H Ni bin tp, nh xut bn nng nghip H Ni nm 2001. Cc tp tip theo (tp 2, tp 3) do Gio s Nguyn Tin Bn l ch bin son. B sch a ra tn gi thc vt c uy tn Vit Nam tham gia bin son. B sch a ra tn gi cc loi thc vt tht chnh xc, bao gm tn gi ca c thc vt bc thp, thc vt bc cao bao gm tt c cc ngnh Vit Nam. Trn y cp n ngha loi cng c hiu nh n v c bn trong nghin cu a dng sinh hc. Bi vy trong nghin cu a dng sinh hc cc ngun ti liu khc nhau v h thc vt, v phn loi thc vt, ni rng ra l v sinh vt l khng th thiu khi cp, khi thc hin nhng nghin cu v a dng sinh hc. Khi m khoa hc k thut cng mi i sng khc nhau ca nhn loi pht trin n nhng nh cao nh ngy nay chng ta ang hng ngy, hng gi chng kin, th cng l lc hng lot cc h sinh thi trn hnh tinh b ph hy, b suy thoi. Hng lot cc loi sinh vt b tuyt chng, v s ng trong s rt nhiu loi ang ng trn b b e da tuyt chng. Khi thut ng a dng sinh hc v khoa hc v a dng sinh hc cng ng thi ra i v ngy cng n hoa kt tri. T c c h thng cc cng trnh, cc ngun ti liu v a dng sinh hc ra i, p ng cc nhu cu nghin cu, nhu cu bo tn a dng sinh hc quy m th gii, quy m quc gia, vng min u c kh y . Vit Nam c nhiu cng trnh nghin cu, nhiu ti liu, sch tra cu v c nhng cun cm nang v a dng sinh hc. C th khi qut rng c ba cp trong nghin cu a dng sinh hc u c th hin. cp phn t c nhng ti liu cp n t khi nim, n cc ni dung cn nghin cu cc phng php nghin cu.iu ny c th hin trong cun Cm nang
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nghin cu a dng sinh vt do nh xut bn Nng nghip pht hnh nm 1997. Hay trong cun Cc phng php nghin cu thc vt cng do tc gi Nguyn Ngha Thn c nh xut bn i hc Quc gia H Ni pht hnh. Cng cp phn t, tc l nhng nghin cu a dng v di truyn, cng nhng nm gn y cng c thm nhng cng trnh nghin cu cng b trn tp ch ca mt s tc gi, m in hnh nht cho hot ng ny l tc gi Nguyn Hong Ngha. Chng hn nh: Phn tch a dng di truyn loi Gii xng (Michelia velutina Candolle, Magnoliaceae H Mc Lan) cho h thc vt Vit Nam ca Nguyn Hong Ngha trong tp ch khoa hc Lm nghip s 3 nm 2009. Hay, phn tch a dng di truyn h gen nhn ca loi M Hi Nam, trong tp ch khoa hc Lm nghip s 2 nm 2009 ca cc tc gi Nguyn Hong Ngha, Trn Thanh Trng, Tin Pht, Nguyn Vn Phng, L Vn Sn, Chu Hong H. Hay, trong cun Bo tn ti nguyn di truyn thc vt rng tc gi Nguyn Hong Ngha cp n cc khi nim v bo tn ti nguyn di truyn, l do, mc tiu cn bo tn, nguyn nhn mt mt a dng di truyn, cc hnh thc bo tn, nh hng nghin cu, cc s liu c bn 10 vn quc gia v cc loi cp bo tn v ti nguyn di truyn. cp c th tc nhng nghin cu v a dng loi, c hng lot cc cng trnh nghin cu, cc ti liu hng dn nghin cu. C th ni cc hot ng trong nghin cu a dng sinh hc v a dng loi l n r nht Vit Nam? T nhng ti liu mang tnh cht hc thut, cc h thng l thuyt, l lun v a dng sinh hc, n nhng cng trnh, nhng bi bo co cng b ca nhiu tc gi, di nhiu gc , nhiu mc , nhiu cch th hin khc nhau v nghin cu a dng loi. c nhiu ti liu mang y tnh hc thut, l lun v a dng sinh hc ca mt s tc gi nh Phm Bnh Quyn trong cun a dng sinh hc. Trong Phm Bnh Quyn cp n hng lot nhng vn , nhng khi nim, nhng thut ng c dng trong nghin cu a dng sinh hc nh: a dng sinh hc l g, nh lng a dng sinh hc, s phn b ca a dng sinh hc, s tuyt chng ca cc loi, nguyn nhn suy thoi a dng sinh hc v bo v a dng sinh hc Vit Nam Hay trong cun Cm nang nghin cu a dng sinh vt Nguyn Ngha Thn cn nu nn nhng phung php nghin cu a dng loi sau khi tp hp cc ngun t liu t cc nc v c nhng t chc bo tn thin nhin, hot ng xung quanh lnh vc a dng sinh hc (IVCN), v c nhng thnh qu ca cc tc gi Vit Nam. Bn cnh cc ti liu di hnh thc cng b l cc tp sch, cn rt nhiu nhng cng b di dng cc bi vit ng ti trong cc tp ch khc nhau nh: Tp ch sinh hc, tp ch khoa hc lm nghip, cc tc gi cp n a dng loi cc khu vc khc nhau Vit
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Nam. Chng hn nh nhng nghin cu v h thc vt vn quc gia Xun Sn ca V Xun Phng trong tp ch sinh hc. hay nhng nghin cu v thnh phn loi cy thn g trn h sinh thi g i thuc thuc huyn C Chi, thnh ph H Ch Minh Ca ng Vn Sn ng trn tp ch khoa gc Lm nghip. Hay cc tc gi Nguyn Duy Chnh, Nng Vn Tip, Trn Vn Tin vi nghin cu thnh phn loi cy g phn b Nam Cam Ly, thuc Cao nguyn Lm Vin, Lm ng ng trn tp ch sinh hc. hay nhng cng b v a dng loi cho h thc vt Vit Nam ca tc gi ngi Vit v tc gi nc ngoi cng b trn tp ch nc ngoi nh nhng nghin cu v bi ba co ca Nguyn Ngha Thn v D. Harder v a dng sinh hc ca h thc vt vng ni cao Fansipan Vit Nam. cp qun th trong nghin cu a dng sinh hc l a dng h sinh thi, phi k n cng trnh Thm thc vt rng Vit Nam ca Thi Vn Tng vi vic phn loi thnh cc kiu thm thc vt v cc u hp thc vt trn t rng Vit Nam. Trong tc gi phn chia mt cch k lng cc kiu thm thc vt, da trn c s kin thc sinh thi chc chn, kt hp vi qu trnh kho cu thc i rt cng phu, cng vi s tng hp t rt nhiu nghin cu v sinh thi hc thm thc vt ca nhiu tc gi c trong nc v nc ngoi nh: Maurand, Schmid, Schenell, Lng Quy, V c Minh, L Vn hi mi kiu thm thc vt tc gi cng cp n cu trc phn tng, n thnh phn loi, n dng sng V dng sng (hay kiu sinh hot) theo cch phn loi ca Raunkiaer, tc gi Thi Vn Trng tm dng h thng dng sng ca Raunkiaer, mt h thng hin nay thng dng trn th gii tin cho vic so snh vi ph dng sng tiu chun. V li cch chia theo kch thc chiu cao l ph hp vi kch thc dng sng ca. Ngoi ra Raunkiaer cng dng c trng ma v hin tng chi c bao hay khng c bao chia thm cc dng ph trong dng sng ca cc cy c chi trn t cng vi cc dng c bit. Dng sng ca cy c chi trn t tc dng sng m trong ma khng thun li cho s sinh trng (ma tuyt vng n i, vng hn i hay ma kh hn vng nhit i), cy khng c nc sinh l dinh dng bnh thng, chi ngn ca cy cao trn mt t. Raunkiaer cn c vo chiu cao ca cy chia thnh bn dng ch yu. - Megaphanerophytes (Mega), cy to c chi trn mt t, trn 30m. - Mesophanerophytes (Meso), cy va c chi trn mt t, t 8 n 30m. - Microphanerophytes (Micro), cy nh c chi trn mt t, t 2 n 8m. - Nanophanerophytes (Nano), cy thp c chi trn mt t, di 2m. Raunkiaer gp nhng cy g c chiu cao trn 8m vo hai dng trn, tc Megaphanerophytes v Mesophanerophytes vo lm mt dng (c khi l nh vy).
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Ngoi bn dng ca cy c chi trn mt t gp c ph bin ti khp cc vng trn tri t, sau ny Raunkiaer cn tm thy cc dng sng cc vng nhit i m v kh hn nh sau: - Lianes phanerophytes (Liano): Cy c chi trn mt t leo qun. y l mt dng sng rt ph bin trong rng cm nhit i. Dng cy leo c th l thn c hay thn g, song d l thn g cng khng cht g, nn phi da vo nhng cy khc vt ln tng lp thch hp vi iu kin sinh thi ca chng. - Epiphytes phanerophytes (Epi): l dng sng ca nhng cy c chi trn t sng nh v sng bm vo cy khc, rt ph bin trong rng m nhit i. Dng ny c th l thn c hay thn g. Nhng loi cy ny khng mc t t ln, m mc ngay trn thn nhng cy g to, cy g nh. Trong dng ny, c loi ph sinh than g, dn dn pht trin ln rt to, m ly thn cy ch v nh si dy thng lng tht ght li, c th lm cho cy ch cht dn. nhng cy bp c trong rng m nhit i l nhng loi cy in hnh cho dng ny. - Phanerophytes herbaces (Her): L dng sng ca cc loi cy c chi trn t thn c, tc dng sng m thn khng c cht g. Trong ma khng thun li, chi ngn vn cao trn mt t ch khng cht ngang mt t nh dng sng c chi ngang mt t. V d cc loi cy tre, cy s, cy chui, cy c khng thn v c nhng loi cy quyt khng thn, khng ho cht trong mu khng thun li u thuc v dng sng ny. - Phanerophytes suculents (Suc.): L dng sng ca cc loi cy c chi trn mt t thn mng. Dng sng ny thng thy cc vng kh hn ti Nam M, Chu Phi. V d nh cc loi cy xng rng, lng long Sau ny Raunkiaer cn dng thm yu t trng ma phn ra mi hai dng sng ph cho bn dng ln ca cc loi cy c chi trn mt t (Phanerophytes): - SMgo: Cy to c chi trn t, thng xanh, khng bao. - SMeo: Cy va c chi trn t, thng xanh, khng bao. - SMio: Cy nh c chi trn t, thng xanh, khng bao. - SNao: Cy thp c chi trn t, thng xanh, khng bao. - SMgC: Cy to c chi trn t, thng xanh, c bao. - SMeC: Cy va c chi trn t, thng xanh, c bao. - SMiC: Cy nh c chi trn t, thng xanh, c bao. - SNaC: Cy thp c chi trn t, thng xanh, c bao. - DMgC: Cy to c chi trn t, rng l, c bao. - DMeC: Cy va c chi trn t, rng l, c bao.
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- DMiC: Cy nh c chi trn t, rng l, c bao. - DNaC: Cy thp c chi trn t, rng l, c bao. Trong khi , khi tng kt cc dng c chi trn t vng nhit i Nam M th Cain cho rng ch c su kiu sau y: MM (Mega v Mesophanerophytes) MI (Microphanerophytes) Ns (Nanophanerophytes) Ep (Epiphytes phanerophytes) Hp (Phanerophytes herbaces) Dng th hai l dng c chi ngang t (Chamephytes), dng sng ny trong ma khng thun li, b phn trn t s ho cht, n ch mt t mt cht, hay ngang mt t. Dng sng th ba l dng c chi mt t (hemicryptophytes), tc l dng sng m trong ma khng thun li, b phn trn t cht c. Dng ny thng c tuyt bao ph trong ma khng thun li. Dng sng th t l dng c chi di t (Crytophytes), tc l dng sng m trong ma khng thun li cc b phn trn t u cht. Chi bn vo nhng b phn nm di t nh c hay gi. Sau Raunkiaer tnh phn trm t l cc dng sng. l c s so snh cc ph dng sng cc vng khc nhau trn tri t. Cc tc gi nghin cu a dng sinh hc Vit Nam cng dng thang nh gi dng sng da trn thang ca Raunkiaer. Nh Nguyn Ngha Thn nu ra trong cun Cc phng php nghin cu thc vt gm cc thang phn chia dng sng nh sau: - Cy chi trn to (Megaphanerophytes): L cc cy g cao t 25m tr ln nh: Sng, Ch ch, Ch xanh, Lim. - Cy chi trn nh (Mesophanerophytes) gm nhng cy g t 8 25m nh cc loi Gi, Sung, Mu ch, Trng. - Cy chi nh (Microphanerophytes) gm nhng cy g nh cao t 2 8m nh cc cy Chi mi, Du da, Ngi, Mn, o. - Cy chi trn ln (Nanophanerophytes) gm nhng cy bi ln v la bi, cy ha g, nh cc loi thuc h C ph, Thu du, R, Gai di tn rng hay cc loi nh Bng bng, Hoa hng, Nhi.
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- Cy b sinh (Epiphytes): gm cc loi cy b sinh sng lu nm trn thn, cnh cy g, trn vch nh cc loi cy trong ngnh dng x, cc loi cy h Phong lan thng sng bm vo cc loi cy g ln trong rng. - Cy k sinh hay bn k sinh (Parasit Hemiparanit phanerophytes) cc loi cy nh Tm gi, T xanh, T hng. - Cy mng nc (Succulentes) nh cc loi cy xng rng, thuc bng. - Dy leo (Lianophanerophytes) gm cc loi cy leo thn c hay thn g cc loi cy nh Kim Ngn, Bm bm, Vng. - Cy chi st t (Chameaphytes) gm nhng cy c chi cch mt t di 25cm nh Cao cng. - Cy chi na n (Hemicryptophytes) gm nhng cy c chi nm st mt t, hay na trn di t. - Cy chi n (Cryptophytes) gm nhng cy c chi nm di t, bao gm cc loi c c, hay cn hnh nh c tranh, gng, c gu, khoai ty - Cy mt nm (Therophytes) gm nhng cy vo thi k kh khn, ton b cy cht i ch cn duy tr ni ging di dng ht. Trn c s phn chia cc dng sng , cc dng sng s c th hin trong cc kt qu nghin cu trong bng danh lc thc vt, sau thng k cc loi theo kiu dng sng ta c ph dng sng di dng cc biu . Trong nghin cu a dng h sinh thi cc tc gi cn c th nh gi nhiu. nhiu l s lng c th ca mt loi no y tiu chun hay mt kiu thm thc vt no y trong mt khu vc no y trong mt khu vc nht nh ty thuc vo mc ch ca vic nghin cu t ra. Ngi ta c th nhiu b sung vo cc bng nghin cu, v d nh nh gi theo ch quan nh mt thang thng k no y. Ngi ta a ra nhiu loi thang thng k nhiu khc nhau v d: Thang nm bc: - Bc 1: Rt t - Bc 2: t - Bc 3: Khng nhiu - Bc 4: Nhiu - Bc 5: Rt nhiu Thang su cp: Thang ny cn c gi l thang Drude, c th hin khng phi bng s m th hin bng ch:
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Soc: S lng c th cy kt li thnh tng kn Cop3: S lng cy rt nhiu Cop2: S lng cy nhiu Cop1: S lng cy tng i nhiu Sp: Thc vt gp vi s lng khng ln lm, tha Sol: Gp rt t tin hnh cc nghin cu a dng h sinh thi, a dng sinh hc cc nh nghin cu thong s dng phng php tiu chun. m t qun x thc vt ring bit no y, cn phi phn chia chng ra cc tiu chun. Cc o tiu chun ny cn phi c trng cho ton b qun x thc vt nhin cu. i vi cc qun x thc vt c, c th din tch dng cho l t 1m2 n 1.000m2, ty thuc vo ln t nhin ca qun x c . i vi qun x thm thc vt rng c th s dng din tch l t 1.00m2 n 1.000m2 hay 5.000m2. ng bng c th t tiu chun ln hn, cc qun x thc vt rng trn ni th din tch ca tiu chun c th nh hn v y iu kin sinh thi phong ph hn v qun x thc vt thay th nhanh. ng thi cc tc gi cng ch ra: Trong cc din tch ny ch m t thm cy g v cy bi, cn thm c di rng th nn chia thnh nhng nh hn, thong th nhng nh hn ny oc gi l ph thuc tiu chun, v kch thc c th l 1m x 1m, 2m x 2m. Cc hnh vung hay ch nht th gc oc lm du bng mt ci cc hay mt du hiu g y. Nu nh din tch m t khng ln lm (thong cho cc qun x thc vt trn ni) c th gii hn tiu chun bng ranh gii t nhin ca qun x . Khng nn a tt c ai chuyn tip ca qun x khc vo. Nu nh ln t nhin cu qun x thc vt ln th c th chia ra thnh cc tiu chun b hn. tiu chun cn oc t nhng ni c trng, c bit l nhng ni khng c iu kin t nhiu. mi qun x khng nn t qu doi ba tiu chun. C th ni rng trong cc ngun ti liu ni v a dng h sinh thi, a dng sinh hc hay cc ti liu v phng php nghin cu thc vt, cc tc gi tng kt cc kinh nghim nghin cu thc vt, nghin cu sinh thi hc ca cc nh sinh thi hc, phn loi hc, a dng sinh hc nhiu ni trn th gii vo bc hng u ch ra cc ch dn, cc quy tc, cc li khuyn cho vic nghin cu a dng sinh hc cc kiu thm thc vt khc nhau, trn nhng vng lnh th c cc iu kin t nhin c trng ring r, khc nhau. iu ny tr nn kho t liu qu gi gip ch chng ti trong qu trnh nghin cu.
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2. Tnh hnh nghin cu a dng thc vt rng thng ba l ( Pinuskesiya) mc t nhin Vit Nam v Lm ng. a) Vit Nam Nh ta bit loi thng ba l (Pinuskesiya) l loi cy g ln, l loi a sng v chu oc hn vo ma kh. Vit Nam loi thng ba l phn b mt s khu vc v to nn kiu rng tha cy l kim a sng chu hn. Thng th kiu rng ny l kiu rng thun loi v thng ba l l loi cy u th sinh thi, chng to nn kiu qun x thc vt rt c trng cho vng ni cao, c bit l Lm ng. Trong cc ngun ti liu khc nhau, loi thng ba l (Pinuskesiya) cn mang nhng tn gi khc l thng nha ngo vi cc tn ng ngha nh: Pinus khasya Royle Pinus langbiangensis Chevalier Thuc h Pinaceae, ngnh Pinophyta. Cng trong cc ngun ti liu khc nhau, loi thng ba l c dn ra cc khu phn b khc nhau Vit Nam. Nh trong tp 1 b cy g rng min Bc Vit Nam do nh xut bn Nng thn pht hnh, c ch ra thng ba l phn b : Lai Chu, Lng Sn, Cao Bng, Qung Ninh v thng mc thnh rng thun loi. Cn tc gi V Vn Chi trong cun Danh lc thc vt Ty Nguyn li nhc ti thng ba l phn b Gia Lai Kom Tum, k Lk, Lm ng. Thc ra loi thng ba l cn phn b mt s khu vc khc Vit Nam nh Bch M, Tha Thin Hu quy m qun th rt nh. Nhn chung thng ba l phn b kh rng Vit Nam vi quy m kch thoc qun th khng ln. Ch ring Lm ng qun th thng rt ln, rt c trng cho kiu thm thc vt y, v rt ng c nghin cu v a dng sinh hc ca kiu rng ny. Trong nhng nghin cu ca mnh, Thi Vn Trng ch ra kiu rng tha cy l kim hi kh nhit i l mt trong ba kiu rng tha. Kiu rng tha hi kh cy l kim nhit i phn b Vit Nam c min Bc v min Nam. min Bc phn b B Trch (Bnh Tr Thin), Hong Mai (Ngh Tnh), Qung Yn (Qung Ninh). min Nam phn b k Lk, Djining, Lt. theo Maurand, qun h thng ba l xung quanh Lt chim mt din tch rng n 70.000 ha. Cng trong cun Thm thc vt rng Vit Nam Thi Vn Trng cho rng u hp thng ba l hay cn gi l u hp ngo. Trong kiu ph min thc vt thn thuc vi khu h di c Himalya Vn Nam Qu Chu. u hp ny chim gn ht vi cc i ni quanh Lt v xung tn pha di trn cc ra ni pha nam cao khong 1.100m, pha ng cao 800m v pha ty trong vng dip thch cao 1.000m. N ln ti cao 1.800m trn
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sn Bono n Lna, v mc kh lin tc ven thung lng sng a Nhim gia dy ni Langbiang v Bidup, sau cng n chim mt phn thng lu sng Krng P, pha ty bc Hou Cris (Schmid, 1962). Rng thng ny bao ph mt din tch rt ln. Ch tnh nguyn khu rng quanh cc i ni xung quanh thnh ph Lt min Nam Vit Nam chim ti 70.000ha. Tng trn ch gm loi ngo hay thng ba l (Pinus kesiya). Mc d loi thng ny l mt loi thng vng cao, cng c ngi cho n t cc trin ni Indonesia trn ln. C ngi khc li cho rng loi thng ba l l loi thng Vn Nam phn ha ra. Nh vy thng ba l (Pinus kesiya) vi kiu rng thng ba l Vit Nam c cc nh thc vt, sinh thi hc rt quan tm nghin cu. ng thi y l loi cy g ln, c th cung cp mt lng g ng k, nn c cc nh nghin cu lm hc quan tm nh mt i tng kinh doanh g c gi tr. Chng th m ngay trong tp 1 ca b cy g rng Vit Nam, Vin quy hoch lm nghip c cng b v loi ny. l cha k n cc hot ng khc nhau ca ngnh lm nghip, cng cc nghin cu ngha ti nguyn lm thuc ca loi ny, ly nha, ly tinh du v cc sn phm khc ly ra t thng ba l. Tuy cha c nghin cu chuyn kho v a dng sinh hc cho rng thng ba l mc t nhin Lm ng, nhng c mt s nghin cu cp s b n thnh phn mt s loi kiu rng ny. Chng hn, cng trong Thm thc vt rng Vit Nam Thi Vn Trng c m t: Tng tn ca rng thng ba l (Pinus khasya) ny gm nhiu loi d vng cao (Quercus.lanata, Quercus helferiana, Lithocarpus dealbathus, Lithocarpus pynostachya) v c nhng loi c hu thy vng Djiring v Lt tc gm nhiu loi trong h Quyn (Ericaceae) nh: cp mc hnh sao (Craibiodendron stellatum), Quyn hoa trng (Pieris ovalifolia). H chua nem (Vacciniaceae) nh: chua nem (Vaccinium exaristatum), Agapetes.sp. Nhng cy h la to thnh tng c nh cc chi: Polytocca, Arundinella, Callipedium, Exotheca, Eulalia, Pogonatherum, Dimeria, Kerriochloa. Cc cy thuc h dng x thng gp nhiu hn c l loi Dicranopteris linearis, Brainia insignis, Woodwardia cochinchinensis, Pteridium aquilinum v Dipteris conjugata. C th ni nhng nghin cu ny l ngun ti liu tham kho ng qu cho nhng nghin cu ca chng ti, khi chnh thc s tr thnh chuyn kho v a dng sinh hc thc vt rng thng ba l Lm ng. b) Lm ng c mt s cng trnh nghin cu v loi thng ba l (Pinus kesiya), v h sinh thi rng thng hay mt s loi cy thuc phn b doi tn rng thng a im ny hay a im kia Lm ng ca mt s tc gi.
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C nhng nghin cu v loi thng ba l theo hng phn loi hc, chng hn nh nghin cu v cc loi cy hc Pinaceae phn b Lm ng ca Nguyn Duy Chnh v Ph c nh c cp n loi thng ba l (Pinus khasya), loi thng nha (Pinus merkusii), loi du sam (Keteluria) mc t nhin Lm ng, cp n danh php, cc kt qu gii phu, c ng trn tm tt bo co ti hi ngh quc t v thc vt t chc ti Nht Bn nm 1993. Trong bo co ti khoa hc cp b ca B gio dc v o to nm 2004, tc gi Lm Ngc Tun vi ti Nghin cu thnh phn cu trc ca h sinh thi rng thng, vai tr ca rng thng i vi mi trng cnh quan Lm ng cp n mt s vn nh: thnh phn cc loi nm rng thng, thnh phn cc loi thc vt bc cao rng thng, thnh phn cc loi chim rng thng, ng vt khng xng sng rng thng, vi sinh vt t rng thng. l nhng ni dung kho st rng ri ca nhiu i tng trong phn loi k c thc vt v ng vt. ng thi cc nghin cu ca tc gi cng cp n mi trng nn mng cho s sng, l t rng thng v sinh thi kiu rng thng Lm ng. Trong nghin cu v sinh thi rng thng Lm ng, tc gi nghin cu c sinh thi rng thng ba l (Pinus khasya) v c sinh thi rng thng hai l (Pinus merkusii) l kiu rng thng tha cy l kim. Phn b ai cao thp, t ph bin nh kiu rng thng ba l. Tc gi cn cp n vai tr ca rng thng, mi quan h ca rng thng v kh hu vai tr ca thin cht lng khng kh ca rng thng. Li c mt s nghin cu cp n a dng sinh hc ca rng thng ba l, tuy cha phi l chuyn kho v kiu rng tha cy l kim ny phn b Lm ng, nhng bc u c mt s kt qu ng tham kho, nhng nghin cu ny l nhng nghin cu nm trong mt d n hay mt tiu ti no . Chng hn nhng nghin cu thuc v tit d n BC trc thuc d n hnh lang a dng sinh hc tiu vng sng M Kng. Trong bo co ca tc gi Nguyn Duy Chnh v Trn Vn Tin cp n 3 tiu chun thc hin ti x a Nhim v a Sa cao 1.500m. Hay trong bo co tng kt ti cp b ca Nguyn Ngc King vi ti Tp on cn trng ph hi rng thng ba l Lm ng, c im v bin php phng chng nm 2000 tc gi tin hnh kho st a dng sinh hc 2 tiu chun Lang Hanh v Ni B. Nhn chung cc nghin cu c mt s kt qu, kha cnh ny hay kha cnh kia, mc ny hay mc kia phn nh phn no nhng c trng cho kiu rng thng ba l, cho a dng sinh hc kiu rng ny. Tuy nhin, nhng nghin cu ch thc hin mt vi im Lm ng, trn mt hay hai cao m loi thng ba l phn b. Nhng nghin cu cha thc hin nhiu a im, cha thc hin ng thi nghin cu ca
25
mnh trn tt c cc ai cao m loi thng ba l (Pinus kesiya) vn phn b t nhin. V vy cc nghin cu cha phn nh y tnh a dng sinh hc thc vt ca rng thng ba l mc t nhin Lm ng. Bi nhn duyn , chng ti ng k ti a dng sinh hc thc vt rng thng ba l (Pinus kesiya) mc t nhin Lm ng v vng ln cn vi hy vng s ng gp phn nh vo cng cuc nghin cu rng thng ni ring v a dng sinh hc nhn chung Vit Nam.
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PHN 2: I TNG V PHNG PHP NGHIN CU I. I TNG NGHIN CU i tng nghin cu ca chng ti trong phm vi ti ny l cc loi cy trong kiu rng thng ba l (Pinus kesiya) mc t nhin Lm ng v vng ln cn. Cc loi cy l cc loi thc vt c mch thuc cc ngnh: Psilophyta, Lycopodiophyta, Equisetophyta, Polypodiophyta, Pinophyta v Magnoliophyta. Rng thng ba l (Pinus kesiya) l kiu rng thng mc t nhin, cc rng thng ny c cu trc ba tng: Tng g ln, tng g nh, tng c - bi, vi cu trc ba tng mi phn nh mt cch t nhin nht v trng thi ca rng thng ba l, v thnh phn loi, v dng sng cng nh nhiu c th loi (tn sut bt gp loi). Rng thng c nghin cu phn b trn tt c cc ai cao t 800m n 2.000m, l cao m thc t loi thng ba l (Pinus kesiya) phn b. II. PHNG PHP NGHIN CU Xc nh kch thc thch hp ca tiu chun xc nh kch thc thch hp ca tiu chun dng cho nghin cu a dng sinh hc kiu rng thng ba l, chng ti s dng phng php xp chng. Cc xp chng c kch thc: 10m x 10m, 15m x 15m, 20m x 20m,30m x 30m, 35m x 35m, 40m x 40m.
40m 35m 30m 25m 20m 15m 10m
Sau thng k s loi tng , t kch thc nh n ln v ta c c tng quan s loi v din tch biu th trn th:
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S loi
im un (x) tng ng vi din tch ln va ca , s loi gn nh khng tng. chnh l kch thc thch hp ca tiu chun dng nghin cu.
45 40 35 30 25 20 15 10 5 0 0
Kch thc
10m 15m 20m 25m 30m 35m 40m x x x x x x x 10m 15m 20m 25m 30m 35m 40m
Xc nh v tr nghin cu Sau khi xc nh c kch thc thch hp ca tiu chun, chng ti thit lp cc nghin cu cc cao khc nhau. Thng mi ai cao c 3 nm su trong cc khi rng thng ba l mc t nhin, trnh xa mp rng loi tr yu t hiu ng vng bin. Cc tiu chun phi phn nh mt cch t nhin v thnh phn loi ca kiu rng ny. Rng thng y phi cu trc ba tng (g ln, g nh, c, bi). xc nh ta (v , kinh ), cao so vi mt bin, hng dc dng my nh v GPS. Xc nh thnh phn loi, dng sng, tnh trng loi, nhiu v ch s a dng xc nh thnh phn loi, v tr phn loi v dng sng chng ti s dng phng php hnh thi so snh; thng qua vic iu tra, thu thp mu vt, s dng ti liu tra cu. nhiu xc nh theo thang Drude, thng qua s xut hin loi trong cc c nghin cu cng tuyn iu tra. Ch s a dng c p dng l ch s giu loi Margalef theo cng thc: DMarg. = ln N
S 1
xc nh s lng c th dng phg php m trc tip cy g ln, g nh, bi. Ring vi cy c th dng 1m x 1m v tr bn gc v chnh gia tiu chun tnh s c th, ri quy ra s c th ca tiu chun chnh. S c th theo cng thc : N= x g ln + y g nh + z cy bi + p c.
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Ch s DMarg. chung cho kiu rng thng ba l mc t nhin Lm ng v vng ln cn c xc nh bng ch s trung bnh cng ca cc :
= xi / n
i =1
29
PHN III: KT QU NGHIN CU I. XC NH KCH THC THCH HP CA TIU CHUN
Xc nh kch thc thch hp ca tiu chun nghin cu a dng thc vt rng thng ba l mc t nhin Lm ng bng phng php lp cc xp chng ln nhau 3 im thuc 3 cao khc nhau, kt qu ch ra : LB01: Ta d a l 120223,7 v bc 1082538,1 kinh ng. cao: 1794m. dc 30, hng dc: TB-N. Biu th tng quan gia kch thc xp chng v s loi:
S loi
Kch thc xp chng 10mx10m 15mx15m 20mx20m 25mx25m 30mx30m 35mx35m 40mx40m
S loi 22 29 34 37 39 40 40
45 40 35 30 25 20 15 10 5 0 0 10 x 15m 20m x 25m x 30m x 35m x 40m x 10m x15m 20m 25m 30m 35m 40m
Kch thc
Nhn xt: Khi kch thc cc chng ca LB01 tng th s loi thc vt c mch cng
tng theo n kch thc 35m x 35m th gn nh khng tng v trn th c mt im un.
TL01: ta a l 115302 v bc, 1082557,0 kinh ng. cao: 1462m.
dc:15. Hng dc: N-B Biu th tng quan gia kch thc xp chng v s loi:
S loi
Kch thc xp chng

10mx10m 15mx15m 20mx20m 25mx25m 30mx30m 35mx35m 40mx40m
S loi 21 28 33 37 40 42 42
45 40 35 30 25 20 15 10 5 0 0
10m x 15m 20m x 25m x 30m x 35m x 40m x 10m x15m 20m 25m 30m 35m 40m
30
Kch thc
Nhn xt: Khi kch thc cc chng ca TL01 tng th s loi thc vt c mch cng tng
theo n kch thc 35m x 35m th gn nh khng tng , v trn th c mt im un

DL01: Ta a l 112320,6 v bc 1080522,0 kinh ng. cao: 890m .
dc: 25. Hng dc: -T Biu th tng quan gia kch thc xp chng v s loi .
S loi
Kch thc xp chng 10mx10m 15mx15m 20mx20m 25mx25m 30mx30m 35mx35m 40mx40m
S loi 20 27 32 38 40 40 41
45 40 35 30 25 20 15 10 5 0 0 10m x 15m x 20m x 25m x 30m x 35m x 40x40 10m 15m 20m 25m 30m 35m
Kch thc
Nhn xt: Khi kch thc cc chng ca DL01 tng th s loi thc vt c mch cng
tng theo n kch thc 35m x 35m th gn nh khng tng v trn th c mt im un. Nh vy t kt qu nghin cu tng quan s loi v kch thc cc xp chng c ba tiu chun (LB01, TL01, DL01) cho bit kch thc thch hp cho vic nghin cu a dng sinh hc thc vt kiu rng thng ba l (Pinus kesiya) mc t nhin Lm ng ca tiu chun l: 35m x 35m.
II. CC KT QU NGHIN CU A DNG SINH HC THC VT CC TIU CHUN.
p dng kch thc tiu chun 35m x 35m cho vic nghin cu a dng thc vt rng thng ba l (Pinus kesiya) mc t nhin tnh Lm ng v vng ln cn. Cc c thit lp trn cc ai cao khc nhau t 800m n 2000m, c dc v hng dc khc
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nhau ch yu trn cc a hnh chia ct kh mnh, cc v tr c xc nh bng v v kinh theo phng php nu trn. ng thi s lng v thnh phn loi, s cc th v ch s a dng Margalef (DMarg) cng c xc nh bng cc phng php v cng thc tnh c nu trn phn phng php nghin cu. Cc kt qu nghin cu c ch ra nh sau:
LB01
V tr: 120 0223,7 v Bc 1080 2538,1 kinh ng cao so vi mt nc bin : 1.794 dc Hng dc S loi (S) S c th (N) - Cy g ln : 22 - Cy g nh : 51 - Cy dng bi : 780 - Cy dng c : 42.875 Tng s c th ca LB01, N = 43.728 DMarg = 3,65 : 300 : TB-N : 40 loi
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Bng: Danh lc thnh phn loi, dng sng v tnh trng loi LB01 TT
Ngnh, h, tn khoa hc loi Ngnh LYCOPDIOPHYTA H Selaginellaceae
Tn vit loi
Dng sng
Tnh trng
1 2 3 4 5 6
Selaginella monospora Spring

H Aspleniaceae
Quyn b n bo t Rng can x gn Rng bit x ui phng l si
Thero. Epi. Nano. Epi.
Asplenium affine Sw.

H Blechnaeae
Brainea insignis ( HooK.) J. Smith

H Polypodiaceae
Crypsinus rhynchoplyllus (HooK.) Copel.

H Polypodiaceae
Pyrrosia lingua (Thunb.) Farw. Var. lingua

H Pteridaceae
Rng tai chut li dao Epi. Seo g, chn x Cryp.
Pteris vittata L.
Ngnh PINOPHYTA H Pinaceae
Pinus kesiya Royle ex Gordon

Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Acanthaceae
Thng ba l
Macro.
8 9 10 11 12 13 14 15 16
Lepidagathis hyalina Ness

H Apriaceae
Ln chy Rau m Anh l petelot. Riu Chn voi nhm Bu t, tam tht gi Nh cc Hong lin r Vt dai
33
Thero. Chamae. Chamae. Thero. Chamae. Chamae. Thero. Nano. Micro.
Centella asiatica (L.)Urb.

H Asteraceae
Ainsliaea petelotii Merr. Colobogyne langbianense Gagn. Elephantopus scaber L. Gynura divaricata (L.) DC. Sonchus oleraceus L.
H Berberidaceae
Mahonia nepalensis DC.

H Caprifoliaceae
Virburnum coriaceum Bl.
H Chloranthaceae
17 18
Chloranthus japonicus Sieb.

H Ericaeae
Si nht Cp mc hnh sao
Thero. Micro.
Craibiodendron stellatum (Pierre ex Lanees) W.W. Sm
19
Vaccinium iteophyllum Hance

H Euphorbiaceae
Nem l liu
Micro.
20 21 22
Aporusa serrate Gagnep.

H Fabaceae
Tai ngh rng Trng qu ngh thut Chm cua
Micro. Nano. Nano.
Desmodium concinnum DC. Indigofera stachyodes Lindl.

H Fagaceae
23 24 25 26 27
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus helferiana A.DC.

H Gentianaceae
D trng Si Helfer. Long m Langbian Cha d Dum Trung b
Micro. Micro. Thero. Nano. Nano.
Gentiana langbianensis A, Chev. ex H. Smith

H Lamiaceae
Elsholtzia blanda (Benth.) Benth.

H Rosaceae
Rubus annamensis Card.

Lp LILIOPSIDA H Asparagaceae
28 29 30 31 32 33 34
Asparagus filicinus Buch Ham ex D.Don.

H Commelinaceae
Thin mn rng Bch trai Burman Mao th trn di Trc tho lng C l gng Xoang trc sc Tc hnh tm
34
Lian. Thero. Thero. Chamae. Cryp. Chamae. Chamae.
Cyanotis papilionacea Roem. & Schult.f.

H Cyperaceae
Fimbristylis oblonga T.Koyama

H Poaceae
Arudinella setosa Trin. Axonopus combressus (Sw.) Beauv. Coelorachis striata (Steud.) A. Cam. Digitaria violascens Link
35 36 37 38 39 40
Eragrostis zeylanica Nees & Mey. Eulalia velutina (Hack.) Kuntze Themeda caudata (Nees.) HacK.
H Smilacaceae
Tinh tho tch lan Ct v lng C phao Kim cang thon Ngi tin Gng
Chamae. Chamae. Chamae. Lian. Cryp. Cryp.
Smilax lanceifolia Roxb.

H Zingiberaceae
Hedychium gardnerianum Rosc. Zingiber rubens Roxb.

LB02
V tr: 120 0227,1 v Bc 1080 2532,9 kinh ng cao so vi mt nc bin : 1.988m dc Hng dc S loi (S) S c th (N) - Cy g ln : 21 - Cy g nh : 47 - Cy dng bi : 712 - Cy dng c : 44.100 Tng s c th ca LB02, N = 44.880 DMarg = 3,55 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi BL02 TT
Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Adiantaceae
: 150 : T- : 39 loi
Tn vit loi
Dng sng
Tnh trng
1 2 3
Adiantum stenochlamys Bak.

H Blechnaeae
Rng nguyt s Rng bch hoa nht Rng vi ln

35
Cryp. Chamae Cryp.
Woodwardia Japonica (L.f.) J.Sm.

H Dennstaedtiaceae
Microlepia platyphylla (D.Don) J.Smith
H Polypodiaceae
Paragramma banaensis (C. Chr). Ching

Rng song vn B N
Epi.

Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Apriaceae
Thng ba l
Macro.
6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22
Pimpenella diversifolia DC.

H Asclepiadaceae
Bng bin
Chamae.
Hoya macrophylla Blume.

H Asteraceae
H da l to C ct heo Bch nhung sa Miu nh Sn cc ni B ch, bp l ng sm Java Vt dai Si nht
Epi. Chamae. Thero. Chamae. Thero. Nano. Lian. Micro. Thero. Micro. Micro. Micro. Micro. Nano. Nano. Nano.
Ageratum conyzoides L. Anaphalis lactea Maxim Hypochaeris radicata L. Wedenia wallichii Lees
H Buddleiaceae
Buddleia asiatica Lour.

H Campanulaceae
Codonopsis Javanica (Bl.) HooK.f.

H Caprifoliaceae

H Chloranthaceae

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Cp mc hnh sao Lyonica ovalifolia (Wall.) Drude Vaccinium iteophyllum Hance
H Euphorbiaceae
Ca di xoan Nem l liu Sc daltonii B ngt hai mu Trng qu ngh thut Tp m l nh

36
Glochidion daltonii (Muel-Arg.) Kurz. Sauropus bicolor Craib

H Fabaceae
Desmodium concinnum DC. Flemingia lincata var. glutinosa Prain
H Fagaceae
23 24 25
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus lanata Smith in Rees.
H Melastomataceae
D trng Si lng Mua lng
Micro. Micro. Nano.
Melastoma saigonense (Kuntze) Merr.

Lp LILIOPSIDA H Amaryllidaceae
26 27 28 29 30 31 32 33 34 35 36 37 38 39
Hypoxis aurea Lour.

H Commelinaceae
H trm, tin mao Bch trai hoang Song bo Mao th trn di H bin ru Xoang trc sc C chn nhn Tinh tho en Ct v lng C tranh C ng C bng lau Th phc linh Gng
Chamae. Thero. Chamae. Thero. Thero. Chamae. Thero. Chamae. Chamae. Cryp. Cryp. Chamae. Lian. Cryp.
Cyanotis vaga (Lour.) Roem. & Schult.f.

H Convallariaceae
Disporum cantonense ( Lour.) Merr.

H Cyperaceae

H Orchidaceae
Habenaria ciliolaris (L.) Kraenzel

H Poaceae
Coelorachis striata (Steud.) A. Cam. Digitaria radicosa (Prese) Miq. Eragrostis nigra Nees ex Steud Eulalia velutina (Hack.) Kuntze Imperata cylindrica (L.) P.Beauv. Panicum repens L. Saccharum spontaneum L.
H Smilacaceae
Smilax glabra wall.ex.Roxb.

H Zingiberaceae
Zingiber rubens Roxb.

LB03
V tr: 120 0228,8 v Bc 1080 2540,8 kinh ng cao so vi mt nc bin : 1.890m dc Hng dc : 120 : TB-N
37
S loi (S) S c th (N) - Cy g ln : 23 - Cy g nh : 72 - Cy dng bi : 954
: 41 loi : 39.024
- Cy dng c : 37.975 Tng s c th ca LB03, N = 39.024 DMarg = 3,78 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi LB03 TT
Ngnh, h, tn khoa hc loi Ngnh LYCOPDIOPHYTA H Selaginellaceae
Tn vit loi
Dng sng
Tnh trng
1 2 3 4 5

H Blechnaeae
Quyn b n bo t Rng bit x Rng i dc Gut, t, rng ty sn Rng tai chut li dao
Thero. Nano. Cryp. Cryp. Epi.

H Dennstaedtiaceae
Pteridium aquilinum (L.) Kuhn.

H Gleicheniaceae
Dicranopteris linearis (Burn.) Underw.

H Polypodiaceae

6 7 8 9 10 11

H Apriaceae
Thng ba l Rau m Riu Thng lo, tai hm Chn voi nhm C chua l
Macro. Chamae. Thero. Thero. Chamae. Thero.

H Asteraceae
Colobogyne langbianense Gagn. Conyza canadense (L.) Crong. Elephantopus scaber L. Emilia sonchifolia (L.) DC
38
12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30
Gerbera piloselloides (L.) Cass. Sonchus oleraceus L.

H Draseraceae
Cc lng, Nh cc Bt rui
Chamae. Thero. Thero. Micro. Micro. Micro. Micro. Micro. Nano. Chamae. Lian. Nano. Micro. Micro. Nano. Chamae. Nano. Chamae. Micro.
Drosera burmannii Vahl.

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Cp mc hnh sao Lyonica ovalifolia (Wall.) Drude
H Euphorbiaceae
Ca di xoan Sc daltonii Me rng Doi Chevalier Bin hng rng thng Trng qu nhiu hoa ng ba tri c cng Chm cua Si kerr Si lng Hng nhu x Gi hc Mua lng An bch sao Du ru
Glochidion daltonii (Muel-Arg.) Kurz. Phyllanthus em blica L

H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Campylotropis pinetorum (Kurz) Schindl. Desmodium multiflorum DC. Dunbaria polocarpa Kurz. Indigofera stachyodes Lindl.
H Fagaceae
Quercus kerri Craib. Quercus lanata Smith in Rees.

H Lamiaceae
Elsholtzia winitiana Craib

H Leeaceae
Leea rubra Bl. ex Spreng.

H Melastomataceae
Melastoma saigonense (Kuntze) Merr. Osbeckia stellata Buch. Ham. ex D.Don

H Myricaceae
Myrica esculenta Buch. Ham. ex D.Don var. Chevalieri (Dode) Pham Hoang
Lp LILIOPSIDA H Commelinaceae
31 32
Cyanotis papilionacea Roem. & Schult.f. Cyperus halpan L.

H Phormiaceae
39
Bch trai Burman C cm
Thero. Chamae.
33 34 35 36 37 38 39 40 41
Dianella ensifolia (L.) DC.

H Poaceae
Hng bi C thm C chn nhn Ct v lng vng K ni X t mnh C phao Kim cang thon Gng
Chamae. Cryp. Thero. Chamae. Thero. Thero. Chamae. Lian. Cryp.
Axonopus affinis A.Chase Digitaria radicosa (Prese) Miq. Eulalia phaeothrix (Hack.) Kuntze Panicum notatum Retz. Sporobolus tenellus Bal. Themeda caudata (Nees.) HacK.
H Smilacaceae
Smilax lanceifolia Roxb.

H Zingiberaceae
TL01
V tr: 110 530,2 v Bc 1080 2557,0 kinh ng cao so vi mt nc bin : 1.462m dc Hng dc S loi (S) S c th (N) - Cy g ln : 22 - Cy g nh : 70 - Cy dng bi : 871 - Cy dng c : 39.200 Tng s c th ca TL01, N = 40.163 DMarg = 3,65 : 150 : N-B : 42 loi
40
Bng: Danh lc thnh phn loi, dng sng v tnh trng loi TL01
TT Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Gleicheniaceae
Tn vit loi
Dng sng
Tnh trng
1 2

H Polypodiaceae
Gut, t, rng ty sn Rng a tc
Cryp. Epi.
Gonophlebium subauriculatum (Bl.) Prest.


Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Asteraceae
Thng ba l
Macro.
4 5 6
Bidens pilosa L. Galingsoga parviflora Cav. Spilanthes paniculata wall. ex DC.

H Ericaeae
n but, Vi cc Cc o, nt o
Thero. Thero. Thero.
7 8
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Cp mc hnh sao Lyonica ovalifolia (Wall.) Drude
H Euphorbiaceae
Micro. Micro.
Ca di xoan
9 10
Aporusa serrate Gagnep. Glochidion daltonii (Muel-Arg.) Kurz.

H Fabaceae
Tai ngh rng Sc daltonii
Micro. Micro.
11 12 13 14 15 16 17
Cassia mimosoides L. Desmodium griffithianum Benth. Desmodium umbellatum (L.) Dc. Dunbaria polocarpa Kurz. Flemingia lincata var. glutinosa Prain Flemingia macophylla (Willd.) Prain Indigofera nigrescens Kurz. ex King & Prain
Mung trinh n Trng qu Griffith. Trng qu tn ng ba tri c cng Tp m l nh Tp m l to Chm en
Thero. Chamae. Thero. Lian. Nano. Nano. Nano.
41
H Fagaceae
18 19 20
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus lanata Smith in Rees. Quercus setulosa Hickel& A. Camus
H Lauraceae
D trng Si lng Si dui
Micro. Micro. Micro.
21 22 23 24 25
Lindera spicata Kosterm.

H Lythraceae
Lin n gi Lun tho l trn Mua chevalier An bch u An bch sao
Micro. Thero. Nano. Thero. Chamae.
Rotala rotundifolia (HooK.f. ex Roxb) Koehne

H Melastomataceae
Melastoma chevalieri Guill. Osbeckia cupulasis D.Don ex W. & Arn. Osbeckia stellata Buch. Ham. ex D.Don
H Myrsinaceae
26 27 28 29 30
Ardisia annamensis Pit.

H Polygalaceae
Cm ngui Trung b Kch nh, du nng Qun hoa cao Mm xi Ngy hng
Nano. Thero. Micro. Nano. Nano.
Polygala paniculata L.
H Proteaceae
Helicia excelsa (Roxb.) Blume

H Rosaceae
Rubus alceaefolius Poir. Rubus cochinchinensis Traht.

H Rubiaceae
31 32 33 34
Hedyotis rudis (Pierre ex Pit.) Phamh.

H Scrophulariaceae
An in nhm ni Ava Giang ni Hoa tm n
Thero. Thero. Micro. Chamae.
Alectra arvenses (Benth.) Merr.

H Theaceae
Ternstroemia japonica (Thunb.) Thunb.

H Violaceae
Viola inconspicua Blume

Lp LILIOPSIDA H Convallariaceae
35
Disporum cantonense ( Lour.) Merr.
Song bo
Chamae.
42
H Phormiaceae
36 37 38 39 40

H Poaceae
Hng bi Xoang trc ng c Song on Lau Ngoi gip
Chamae. Chamae. Chamae. Chamae. Chamae.
Coelorachis pratensis (Bal.) A.Cam. Diectomis fastigiata (Sw.) Kunth Erianthus arundinaceus (Retz.) Jeswiet. ex Heyne Exotheca abyssinica (A. Rich.) A nders.
H Smilacaceae
41 42

H Zingiberaceae
Th phc linh Gng
Lian. Cryp.

TL02
V tr: 110 5252,2 v Bc 1080 2558,3 kinh ng cao so vi mt nc bin : 1.553m dc Hng dc S loi (S) S c th (N) - Cy g ln : 21 - Cy g nh : 85 - Cy dng bi : 587 - Cy dng c : 34.300 Tng s c th ca TL02, N = 34.993 DMarg = 3,63 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi TL02
TT Ngnh, h, tn khoa hc loi Ngnh PINOPHYTA H Pinaceae
: 320 : TN-B : 39 loi
Tn vit loi
Dng sng
Tnh trng
Thng ba l
Macro.
43
2 3
Eupatorium odoratum L. Gynura crepidoides Benth.

H Campanulaceae
C Lo, c cng sn Rau tu bay
Thero. Thero.
4 5 6 7

H Chloranthaceae
ng sm Java Si nht
Lian. Thero. Micro. Micro.

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Cp mc hnh sao Vaccinium iteophyllum Hance
H Euphorbiaceae
Nem l liu
8 9

H Fabaceae
Sc daltonii Me rng
Micro. Micro.
10 11 12 13 14 15
Campylotropis pinetorum (Kurz) Schindl. Clitoria macrophylla Wall. ex Benth. Crotalaria albida Heyne ex Roth. Flemingia macophylla (Willd.) Prain Indigofera stachyodes Lindl. Shuteria suffutta Benth.
H Fagaceae
Bin hng rng thng Bic l to Sc sc trng Tp m l to Chm cua Mang sang
Nano. Lian. Chamae. Nano. Nano. Thero.
16 17 18
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus helferiana A.DC. Quercus lanata Smith in Rees.
H Juglandaceae
D trng Si Helfer. Si lng
19 20
Engelhardia spicata Lesch. ex Blume

H Leeaceae
Cho bng Gi hc
Micro. Chamae.

H Melastomataceae
44
21 22
Melastoma saigonense (Kuntze) Merr. Osbeckia chinenensis L.

H Myricaceae
Mua lng An bch Trung Quc
Nano. Chamae.
23
H Myrsinaceae
Du ru
Micro.
24 25 26 27 28 29 30 31
Ardisia mirabilis Pit.

H Oxalidaceae
Cm ngui l Me t nh Kch nh, du nng Qun hoa cao Dum Trung b An in tai Ba chc An tc
Nano. Chamae. Thero. Micro. Nano. Thero. Nano. Micro.
Oxalis corniculata L
H Polygalaceae
H Proteaceae

H Rosaceae

H Rubiaceae
Hedyotis auricularia L.
H Rutaceae
Glycosmis pentaphylla (Retz.) DC.

H Styracaceae
Styrax benjoin Dryand

32 33 34 35 36 37 38
Commelina commuris L.
H Orchidaceae
Trai thng Ha hong Hng bi Xoang trc ng c Xoang trc sc C chn nhn Mao phng chm
45
Thero. Epi. Chamae. Chamae. Chamae. Thero. Chamae.
Dendrobium christyanum Reichb.f.

H Phormiaceae

H Poaceae
Coelorachis pratensis (Bal.) A.Cam. Coelorachis striata (Steud.) A. Cam. Digitaria radicosa (Prese) Miq. Erianthus fastigiatus Nees ex Steud
H Zingiberaceae
39

TL03
Gng
Cryp.
V tr: 110 5242,3 v Bc 1080 2610,2 kinh ng cao so vi mt nc bin : 1.383m dc Hng dc S loi (S) S c th (N) - Cy g ln : 25 - Cy g nh : 122 - Cy dng bi : 825 - Cy dng c : 50.225 Tng s c th ca TL03, N = 51.197 DMarg = 3,69 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi TL03
TT Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Gleicheniaceae
: 100 : TN-B : 41 loi
Tn vit loi
Dng sng
Tnh trng

Gut, t, rng ty sn
Cryp.
2 3
Keteleeria evelyniana Mast Pinus kesiya Royle ex Gordon

Du sam ni t Thng ba l
Macro. Macro.
4 5
Dichrocephala integrifolia (L.f.) Kuntre Gynura crepidoides Benth.
Lng st l nguyn Rau tu bay
Thero. Thero.
46
H Euphorbiaceae
6 7 8 9 10 11 12 13 14 15 16 17
Phyllanthus amarus Schum. & Thonn.

H Fabaceae
Dip h chu trng Doi Chevalier Bin hng rng thng Mung trinh n Trng qu tn ng ba tri c cng Tp m l nh Tp m l to Chm en Chm cua on kim ta Hu v hoe
Thero. Micro. Nano. Thero. Thero. Lian. Nano. Nano. Nano. Nano. Chamae. Chamae.
Archidendron chevalieri (Kost.) I. Niels. Campylotropis pinetorum (Kurz) Schindl. Cassia mimosoides L. Desmodium umbellatum (L.) Dc. Dunbaria polocarpa Kurz. Flemingia lincata var. glutinosa Prain Flemingia macophylla (Willd.) Prain Indigofera nigrescens Kurz. ex King & Prain Indigofera stachyodes Lindl. Tephrosia purpurea (L.) Pers. Uraria rufescens (DC.) Schindl.
H Fagaceae
18 19 20 21 22
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Lithocarpus parvulus (Hickel & A.Camus) A.Camus Quercus helferiana A.DC. Quercus kerri Craib. Quercus lanata Smith in Rees.
D trng D gi Si Helfer. Si kerr Si lng
Micro. Micro. Micro. Micro. Micro.
NV01
V tr: 110 5221,4 v Bc 1080 2616,3 kinh ng cao so vi mt nc bin : 1.334m dc Hng dc S loi (S) : 150 : N-B : 43 loi
47
S c th (N) - Cy g ln : 18 - Cy g nh : 65 - Cy dng bi : 982 - Cy dng c : 51.450 Tng s c th ca NV01, N = 52.515 DMarg = 3,86 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi NV01 TT
Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Gleicheniaceae
Tn vit loi
Dng sng
Tnh trng
1 2 3 4 5

Ngnh PINOPHYTA
Gut, t, rng ty sn Thng ba l n but, C chua l Khc vng
Cryp. Macro. Thero. Thero. Thero.

H Asteraceae
Bidens pilosa L. Emilia sonchifolia (L.) DC Gnaphalium luteo-album L.

H Fabaceae
6 7 8 9 10 11 12 13
Desmodium umbellatum (L.) Dc. Dunbaria polocarpa Kurz. Flemingia lincata var. glutinosa Prain Indigofera longicandata Thun Indigofera stachyodes Lindl. Shuteria suffutta Benth. Tephrosia purpurea (L.) Pers. Uraria rufescens (DC.) Schindl.
H Fagaceae
Trng qu tn ng ba tri c cng Tp m l nh Chm ui di Chm cua Mang sang on kim ta Hu v hoe
Thero. Lian. Nano. Nano. Nano. Thero. Chamae. Chamae.
14
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd.
D trng
Micro.
48
15 16 17
Quercus helferiana A.DC. Quercus kerri Craib. Quercus lanata Smith in Rees.
H Juglandaceae
Si Helfer. Si kerr Si lng
18 19 20 21

H Lauraceae
Cho bng Lin n gi Mua lng An bch Trung Quc
Micro. Micro. Nano. Chamae.

H Melastomataceae

H Myrsinaceae
22 23
Ardisia annamensis Pit. Ardisia mirabilis Pit.

H Nepenthacece
Cm ngui Trung b Cm ngui l
Nano. Nano.
24 25 26 27
Nepenthes annamensis Macfarl.

H Proteaceae
Bnh nc Trung b Qun hoa cao Mm xi Ngy hng
Chamae. Micro. Nano. Nano.

H Rosaceae

H Rubiaceae
28 29 30 31 32 33
H Rutaceae
An in tai Gii lm, du da ni Ava kn Con chut Giang ni
Thero. Nano. Thero. Nano. Nano. Micro.
Clausena excavata Burm.f.

H Scrophulariaceae

H Sterculiaceae
Helicteres angustifolia L. Helicteres hirsuta Lour. Ternstroemia japonica (Thunb.) Thunb.

Lp LILIOPSIDA H Cyperaceae
49
34
Kyllinga melanosperma Nees

H Phormiaceae
Bch u
Chamae.
35 36 37 38 39 40 41

H Poaceae
Hng bi Trc tho lng Xoang trc sc Ct v ra San trn Hng nhung L sy
Chamae. Chamae. Chamae. Chamae. Chamae. Thero. Chamae.
Arudinella setosa Trin. Coelorachis striata (Steud.) A. Cam. Eulalia fimbriata (Hack.) Kuntze Paspalum orbiculare Forst. Rhynchelytrum repens (Willd.) C.E. Hubb. Themeda arundinacea (Roxb.) HacK.
H Smilacaceae
42 43

H Zingiberaceae
Th phc linh Gng
Lian. Cryp.
DTL01
V tr: 110 5425,5 v Bc 1080 2728,9 kinh ng cao so vi mt nc bin : 1.228m dc Hng dc S loi (S) S c th (N) - Cy g ln : 21 - Cy g nh : 58 - Cy dng bi : 751 - Cy dng c : 47.775 Tng s c th ca DTL01, N = 48.605 DMarg = 3,80 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DTL01 : 280 : N-B : 42 loi
50
TT
Ngnh, h, tn khoa hc loi Ngnh LYCOPDIOPHYTA H Lycopodiaceae
Tn vit loi
Dng sng
Tnh trng
Lycopodionella cernuua (L.) Pic.Serm.

Ngnh POLYPODIOPHYTA H Adiantaceae
Thch tng nghin
Thero.
2 3 4 5 6

H Aspleniaceae
Rng nguyt s Rng can x gn Rng bch hoa nht Gut, t, rng ty sn Rng vy c nc
Cryp. Epi. Chamae Cryp. Epi.

H Blechnaeae

H Gleicheniaceae

H Polypodiaceae Lemmaplyllum carnosum (HooK.) C. Presl. Ngnh PINOPHYTA H Pinaceae
7 8 9 10 11 12 13

H Anacardiaceae
Thng ba l Mui, Ng bi t H da l to C ct heo Thng lo, tai hm Bch u Sn cc ni
Macro. Micro. Epi. Chamae. Thero. Chamae. Thero.
Rhus chinensis Muell.

H Asclepiadaceae
Hoya macrophylla Blume.

H Asteraceae
Ageratum conyzoides L. Conyza canadense (L.) Crong. Vernonia squarrosa (D.Don) Less. Wedenia wallichii Lees
H Buddleiaceae
14 15 16

H Chloranthaceae
B ch, bp l Si nht Cp mc hnh sao

51
Nano. Thero. Micro.

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees)
17 18

H Fabaceae
Sc daltonii Me rng
Micro. Micro.
19 20 21 22 23
Archidendron chevalieri (Kost.) I. Niels. Crotalaria albida Heyne ex Roth. Desmodium griffithianum Benth. Desmodium sp. Dunbaria polocarpa Kurz.
H Fagaceae
Doi Chevalier Sc sc trng Trng qu Griffith. u l nh ng ba tri c cng
Micro. Chamae. Chamae. Thero. Lian.
24

D trng
Micro.
25 26 27 28 29 30 31
Quercus helferiana A.DC.

H Juglandaceae
Si Helfer. Cho bng Hng nhu x Sc mo ct An bch u Ngi kh Du ru
Micro. Micro. Nano. Thero. Thero. Nano. Micro.

H Lamiaceae
Elsholtzia winitiana Craib

H Loganiaceae
Mitrasaeme eriophila Leenh.

H Melastomataceae
Osbeckia cupulasis D.Don ex W. & Arn.

H Moraceae
Ficus hirta Vahl var. roxburghii (Miq.) King

H Myricaceae
H Myrsinaceae
32 33 34
Ardisia annamensis Pit.

H Nepenthacece
Cm ngui Trung b Bnh nc Trung b Dum Trung b
Nano. Chamae. Nano.

H Rosaceae

H Rubiaceae
52
35
H Commelinaceae
An in tai
Thero.
36 37 38 39 40 41
Murdannia simplex (Vahl.) Brenan

H Cyperaceae
Trai l hp Mao th mnh Trc tho lng Xoang trc sc ui chn nh L nhn
Thero. Thero. Chamae. Chamae. Chamae. Thero.
Fimbristylis gracilenta Hance

H Poaceae
Arudinella setosa Trin. Coelorachis striata (Steud.) A. Cam. Setaria pumila (Poir.) Roem. & Schult. Themeda arguens (L.) HacK
H Smilacaceae
42

DTL02
Th phc linh
Lian.
V tr: 110 5417,4 v Bc 1080 2729,3 kinh ng cao so vi mt nc bin : 1.310m dc Hng dc S loi (S) S c th (N) - Cy g ln : 22 - Cy g nh : 71 - Cy dng bi : 684 - Cy dng c : 39.200 Tng s c th ca DTL02, N = 39.977 DMarg = 3,59 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DTL02
TT Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Adiantaceae
: 350 : N-TB : 39 loi
Tn vit loi
Dng sng
Tnh trng

53
Rng nguyt s
Cryp.
H Aspleniaceae
2 3 4 5 6

H Dennstaedtiaceae
Rng can x gn Rng i dc Song dc i Gut, t, rng ty sn Rng song vn B N
Epi. Cryp. Cryp. Cryp. Epi.

H Dipteridaceae
Dipteris conjugata (Kaulf.) Reinw.

H Gleicheniaceae
Dicranopteris linearis (Burn.) Underw. Paragramma banaensis (C. Chr). Ching


Thng ba l
Macro.
8 9 10 11
Conyza canadense (L.) Crong. Eupatorium odoratum L. Vernonia squarrosa (D.Don) Less.
H Boraginaceae
Thng lo, tai hm C Lo, c cng sn Bch u Khuyn thit
Thero. Thero. Chamae. Thero.
Cynoglossum zeylanicum (Vahl. ex Hornem.) Thunb. ex Lehm.

H Euphorbiaceae
12 13 14 15 16 17 18 19
Aporusa serrate Gagnep. Glochidion daltonii (Muel-Arg.) Kurz. Phyllanthus em blica L

H Fabaceae
Tai ngh rng Sc daltonii Me rng Doi Chevalier Sc sc trng Bch thit An bch Nepal Du ru
Micro. Micro. Micro. Micro. Chamae. Thero. Thero. Micro.
Archidendron chevalieri (Kost.) I. Niels. Crotalaria albida Heyne ex Roth.

H Lamiaceae
Leucas ciliata Benth.

H Melastomataceae
Osbeckia nepalensis HooK.

H Myricaceae
Myrica esculenta Buch. Ham. ex D.Don var.
54
Chevalieri (Dode) Pham Hoang

H Nepenthacece
20 21 22 23 24 25 26 27 28 29 30 31

H Polygalaceae
Bnh nc Trung b Kch nh, du nng Qun hoa cao Ngy hng An in tai xoan ni Ava S bu ch ba kn Chn tr Harman C ke lng Gai u lng
Chamae. Thero. Micro. Nano. Thero. Nana. Thero. Thero. Nano. Nano. Nano. Nano.
H Proteaceae

H Rosaceae
Rubus cochinchinensis Traht.

H Rubiaceae
H Rutaceae
Euodia lepta (Spreng.) Merr.

H Scrophulariaceae
Alectra arvenses (Benth.) Merr. Sopubia trifida Buch. Ham ex G.Don

H Sterculiaceae
Helicteres angustifolia L.
H Theaceae
Eurya Japonica var. harmandii Pierre ex Pitard

H Tiliaceae
Grewia hirsuta Vahl. Triumfetta pseudocana Sprague & Craib.

32 33 34 35 36 37 38

H Cyperaceae
Bch trai hoang C cm
Thero. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae.
Cyperus halpan L.
H Poaceae
Arudinella setosa Trin. Coelorachis striata (Steud.) A. Cam. Eragrostis nutans (Retz.) Nees ex Steud Exotheca abyssinica (A. Rich.) A nders. Setaria pumila (Poir.) Roem. & Schult.
55
Trc tho lng Xoang trc sc Tinh tho nghin Ngoi gip ui chn nh
39
Themeda arguens (L.) HacK

TN01
L nhn
Thero.
V tr: 110 5559,8 v Bc 1080 2231,6 kinh ng cao so vi mt nc bin : 1.378m dc Hng dc S loi (S) S c th (N) - Cy g ln : 19 - Cy g nh : 82 - Cy dng bi : 850 - Cy dng c : 52.675 Tng s c th ca TN01, N = 53.626 DMarg = 3,58 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi TN01
TT Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Polypodiaceae
: 150 :TN-B : 40 loi
Tn vit loi
Dng sng
Tnh trng
1 2
Crypsinus rhynchoplyllus (HooK.) Copel.

H Pteridaceae
Rng n thy c mi Seo g, chn x
Epi. Cryp.
Pteris vittata L.
3 4

H Anacardiaceae
Thng ba l Mui, Ng bi t
Macro. Micro.

Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Apriaceae
Hydrocotyle chevalieri (Chern) Tard.
Rau m chevalier
Chamae
56
H Asclepiadaceae
6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27
Streptocaulon juventas (Lour.) Merr.

H Asteraceae
H th trng n but, i bi l ln Khc vng Sn cc ni B ch, bp l ng sm Java Dy chiu Cp mc hnh sao Nem l liu Sc daltonii Me rng Trng qu tc vng u sam u ba thy D trng Si Helfer. Si dui Bch thit Mua lng An bch Trung Quc Du ru
Lian. Thero. Thero. Thero. Thero. Nano. Lian. Lian. Micro. Micro. Micro. Micro. Thero. Lian. Lian. Micro. Micro. Micro. Thero. Nano. Chamae. Micro.
Bidens pilosa L. Blumea sinuata (Lour.) Merr. Gnaphalium luteo-album L. Wedenia wallichii Lees
H Buddleiaceae

H Campanulaceae

H Dilleniaceae
Tetracera scandens (L.)Merr

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Vaccinium iteophyllum Hance

H Euphorbiaceae

H Fabaceae
Desmodium auricomum Grah. ex Benth. Dunbaria villosa (Thunb.) Makino. Vigna triloba (L.) Ver dc.
H Fagaceae
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus helferiana A.DC. Quercus setulosa Hickel& A. Camus
H Lamiaceae
Leucas ciliata Benth.

H Melastomataceae

H Myricaceae

57

H Oxalidaceae
28 29 30 31 32 33 34 35 36 37 38 39 40
H Rosaceae
Me t nh Mm xi An in tai Ba chc ni Ava Ng sc Bch trai hoang Kit Trc tho lng Xoang trc sc Lau C tranh K ni
Chamae. Nano. Thero. Nano. Thero. Nano. Thero. Chamae. Chamae. Chamae. Chamae. Cryp. Thero.
Rubus alceaefolius Poir.

H Rubiaceae
H Rutaceae

H Scrophulariaceae

H Verbenanaceae
Lantana camara L.
H Commelinaceae

H Cyperaceae
Carex lindleyana Nees

H Poaceae
Arudinella setosa Trin. Coelorachis striata (Steud.) A. Cam. Erianthus arundinaceus (Retz.) Jeswiet. ex Heyne Imperata cylindrica (L.) P.Beauv. Panicum notatum Retz.
SV01
V tr: 110 5918,5 v Bc 1080 2150,4 kinh ng cao so vi mt nc bin : 1.427m dc Hng dc S loi (S) S c th (N) - Cy g ln : 17 - Cy g nh : 65
58
: 350 : N-B : 43 loi
- Cy dng bi : 925 - Cy dng c : 42.875 Tng s c th ca SV01, N = 43.882 DMarg = 3,93 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi SV01
TT Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Blechnaeae
Tn vit loi
Dng sng
Tnh trng
1 2

H Gleicheniaceae
Rng bch hoa nht Gut, t, rng ty sn
Chamae Cryp.


Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Asteraceae Hypochaeris radicata L.
Thng ba l
Macro.
4 5 6 7 8 9 10 11 12 14 15 16
Miu nh Inugn Bch u Si nht Cp mc hnh sao Nem l liu Me rng Doi Chevalier Mung trinh n Trng qu nhiu hoa Thc lp Schubert Trng qu tn
59
Chamae. Thero. Chamae. Thero. Micro. Micro. Micro. Micro. Thero. Chamae. Nano. Thero.
Inula nervosa Wall.ex DC. Vernonia squarrosa (D.Don) Less.

H Chloranthaceae

H Ericaeae

H Euphorbiaceae
Phyllanthus em blica L
H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Cassia mimosoides L. Desmodium multiflorum DC. Desmodium schubertiae Ohashi Desmodium umbellatum (L.) Dc.
17 18 19 20 21 22 23 24 25 26
Indigofera stachyodes Lindl. Tephrosia purpurea (L.) Pers.

H Fagaceae
Chm cua on kim ta D trng Si lng Long m Gi hc Lun tho l trn An bch Trung Quc An bch sao Du ru
Nano. Chamae. Micro. Micro. Thero. Chamae. Thero. Chamae. Chamae. Micro.
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus lanata Smith in Rees.
H Gentianaceae
Gentiana indica Steud.

H Leeaceae

H Lythraceae
Rotala rotundifolia (HooK.f. ex Roxb) Koehne

H Melastomataceae
Osbeckia chinenensis L. Osbeckia stellata Buch. Ham. ex D.Don

H Myricaceae
H Proteaceae
27 28 29 30 31 32 33
Helicia excelsa (Roxb.) Blume Rubus annamensis Card.

H Rubiaceae
Qun hoa cao Dum Trung b An in tai Ba chc kn Giang ni T chu
Micro. Nano. Thero. Nano. Nano. Micro. Nano.
H Rutaceae

H Sterculiaceae
H Theaceae

H Verbenanaceae
Callicarpa rubella Lindl.

34 35
Cyanotis papilionacea Roem. & Schult.f. Murdannia giganteum ( Vahl.) Bruckner

60
Bch trai Burman Trai cao
Thero. Thero.
H Phormiaceae
36 37 38 39 40 41 42 43

H Poaceae
Hng bi Trc tho lng Xoang trc sc C chn nhn Tinh tho tch lan Sao urville Th phc linh Gng
Chamae. Chamae. Chamae. Thero. Chamae. Chamae. Lian. Cryp.
Arudinella setosa Trin. Coelorachis striata (Steud.) A. Cam. Digitaria radicosa (Prese) Miq. Eragrostis zeylanica Nees & Mey. Paspalum urvillei Steud.
H Smilacaceae

H Zingiberaceae

SV02
V tr: 110 5914,6 v Bc 1080 2202,0 kinh ng cao so vi mt nc bin : 1.455m dc Hng dc S loi (S) S c th (N) - Cy g ln : 18 - Cy g nh : 83 - Cy dng bi : 875 - Cy dng c : 47.775 Tng s c th ca SV02, N = 48.751 DMarg = 3,71 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi SV02
: 250 : N-TB : 41 loi
Tn vit loi
Dng sng
Tnh trng
Keteleeria evelyniana Mast
Du sam ni t
Macro.
61

Thng ba l
Macro.
3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24
Elephantopus mollis H.B.K Eupatorium odoratum L.

H Draseraceae
Chn voi mm, cao C Lo, c cng sn Trng l bn nguyt Cp mc hnh sao Ca di xoan Nem l liu Tai ngh rng Sc daltonii B ngt hai mu Doi Chevalier Bin hng rng thng Mung trinh n Trng qu nhiu hoa ng ba tri c cng Tp m l nh Chm cua u ma on kim ta D trng Long m Cho bng Lin n gi
62
Chamae. Thero. Thero. Micro. Micro. Micro. Micro. Micro. Nano. Micro. Nano. Thero. Chamae. Lian. Nano. Nano. Lian. Chamae. Micro. Thero. Micro. Micro.
Drosera peltata Smith in Willd.

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Lyonica ovalifolia (Wall.) Drude Vaccinium iteophyllum Hance
H Euphorbiaceae
Aporusa serrate Gagnep. Glochidion daltonii (Muel-Arg.) Kurz. Sauropus bicolor Craib
H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Campylotropis pinetorum (Kurz) Schindl. Cassia mimosoides L. Desmodium multiflorum DC. Dunbaria polocarpa Kurz. Flemingia lincata var. glutinosa Prain Indigofera stachyodes Lindl. Pueraria phaseoloides (Roxb.) Benth. Tephrosia purpurea (L.) Pers.
H Fagaceae

H Gentianaceae

H Juglandaceae

H Lauraceae
H Melastomataceae
25 26
Melastoma candidum D.Don Osbeckia cupulasis D.Don ex W. & Arn.

H Myricaceae
Mua trng An bch u
Nano. Thero.
27
H Myrsinaceae
Du ru
Micro.
28 29
Ardisia annamensis Pit. Maesa membranacea A.DC.

H Proteaceae
Cm ngui Trung b n mng
Nano. Nano.
30 31 32 33 34 35

H Rosaceae
Qun hoa cao Ngy hng An in nhm Xoan ni Ava Giang ni
Micro. Nano. Thero. Nana. Thero. Micro.

H Rubiaceae

H Rutaceae

H Scrophulariaceae

H Theaceae

Lp LILIOPSIDA H Poaceae
36 37 38 39 40
Arudinella setosa Trin. Axonopus affinis A.Chase Eragrostis nutans (Retz.) Nees ex Steud Imperata cylindrica (L.) P.Beauv. Miscanthus floridulus (Labill.) Warb. ex Schum & Laut.
H Smilacaceae
Trc tho lng C thm Tinh tho nghin C tranh L sng
Chamae. Cryp. Chamae. Cryp. Chamae.
41
Smilax corbularia Kunth.
Kim cang lin hng
Lian.
63
DS01
V tr: 120 0046,8 v Bc 1080 2912,5 kinh ng cao so vi mt nc bin : 1.459m dc Hng dc S loi (S) S c th (N) - Cy g ln : 19 - Cy g nh : 65 - Cy dng bi : 786 - Cy dng c : Tng s c th ca DS01, N = 46.195 DMarg = 3,82 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DS01
TT Ngnh, h, tn khoa hc loi Ngnh POLYPODIOPHYTA H Blechnaeae
: 400 : ,N-T,TB : 42 loi
Tn vit loi
Dng sng
Tnh trng
1 2 3

H Gleicheniaceae
Rng bch hoa nht Gut, t, rng ty sn Rng t minh qu
Chamae Cryp. Cryp.

H Osmundaceae
Osmunda cinnamomea L.

Thng ba l
Macro.
5 6
Colobogyne langbianense Gagn. Gnaphalium luteo-album L.
Riu Khc vng
Thero. Thero.
64
7 8 9 10 11 12 13 14 14 15 17 18 19 20 21 22 23 24 25 26 27 28 29
Inula nervosa Wall.ex DC.

H Ericaeae
Inugn Cp mc hnh sao Nem l liu D fleury Me rng B ngt hai mu Doi Chevalier Bin hng rng thng Trng qu tn u sam Tp m l to u ma D trng Si Helfer. Si lng Long m Cm thy trung vit Lin n gi Mua lng An bch Nepal Cm ngui rng Kch nh, du nng An in nhm
65
Thero. Micro. Micro. Nano. Micro. Nano. Micro. Nano. Thero. Lian. Nano. Lian. Micro. Micro. Micro. Thero. Thero. Micro. Nano. Thero. Nano. Thero. Thero.

H Euphorbiaceae
Breynia fleuryi Beille Phyllanthus em blica L Sauropus bicolor Craib

H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Campylotropis pinetorum (Kurz) Schindl. Desmodium umbellatum (L.) Dc. Dunbaria villosa (Thunb.) Makino. Flemingia macophylla (Willd.) Prain Pueraria phaseoloides (Roxb.) Benth.
H Fagaceae
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus helferiana A.DC. Quercus lanata Smith in Rees.
H Gentianaceae

H Lamiaceae
Nosema cochinchinensis (Lour.) Merr.

H Lauraceae

H Melastomataceae
Melastoma saigonense (Kuntze) Merr. Osbeckia nepalensis HooK.

H Myrsinaceae
Ardisia crenta Sims.

H Polygalaceae
H Rubiaceae
H Rutaceae
30 31 32 33

H Scrophulariaceae
Gii lm, du da ni Ava kn Giang ni
Nano. Thero. Nano. Micro.

H Sterculiaceae
H Theaceae

34 35 36 37 38 39 40 41 42
Murdannia simplex (Vahl.) Brenan

H Cyperaceae
Trai l hp Bch u Trc tho lng C l gng Song on Tinh tho en San trn Bc nh Gng
Thero. Chamae. Chamae. Cryp. Chamae. Chamae. Chamae. Thero. Cryp.
Kyllinga melanosperma Nees

H Poaceae
Arudinella setosa Trin. Axonopus combressus (Sw.) Beauv. Diectomis fastigiata (Sw.) Kunth Eragrostis nigra Nees ex Steud Paspalum orbiculare Forst. Saceiolepis indica (L.) Chase
H Zingiberaceae
DS02
V tr: 120 0045,7 v Bc 1080 2917,4 kinh ng cao so vi mt nc bin : 1.517m dc Hng dc S loi (S) S c th (N) - Cy g ln : 16 - Cy g nh : 28
66
: 250 : TN-B : 39 loi
- Cy dng bi : 515 - Cy dng c : 35.525 Tng s c th ca DS02, N = 36.084 DMarg = 3,62 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DS02
Tn vit loi
Dng sng
Tnh trng

Thng ba l
Macro.
2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17
Galingsoga parviflora Cav. Gynura crepidoides Benth. Wedenia urticaefolia (Blume) DC. ex Wight
H Draseraceae
Vi cc Rau tu bay Sn cc nhm Trng l bn nguyt Ca di xoan Tai ngh rng Sc daltonii Doi Chevalier Gip qu Mung trinh n Sc sc trng Thc lp Schubert ng ba tri c cng Tp m l nh Chm cua Mc c
Thero. Thero. Chamae. Thero. Micro. Micro. Micro. Micro. Lian. Thero. Chamae. Nano. Lian. Nano. Nano. Chamae.
Drosera peltata Smith in Willd.

H Ericaeae
Lyonica ovalifolia (Wall.) Drude

H Euphorbiaceae

H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Cajanus elongatus (Benth) Maesen Cassia mimosoides L. Crotalaria albida Heyne ex Roth. Desmodium schubertiae Ohashi Dunbaria polocarpa Kurz. Flemingia lincata var. glutinosa Prain Indigofera stachyodes Lindl. Mimosa pudica L.
H Fagaceae
67
18 19 20 21 22 23 24 25 26 27 28 29 30 31 32
Quercus setulosa Hickel& A. Camus

H Lamiaceae
Si dui Cau phong lun Hng nhu x Bch thit Gi hc Mua lng Cm ngui Trung b Cm ngui l Me t nh Qun hoa cao Mm xi Ngy hng Xoan S bu ch ba Ng sc
Micro. Thero. Nano. Thero. Chamae. Nano. Nano. Nano. Chamae. Micro. Nano. Nano. Nana. Thero. Nano.
Clinopodium gracile (Benth.) Matsum. Elsholtzia winitiana Craib Leucas ciliata Benth.
H Leeaceae

H Melastomataceae
Melastoma saigonense (Kuntze) Merr.

H Myrsinaceae
Ardisia annamensis Pit. Ardisia mirabilis Pit.

H Oxalidaceae
H Proteaceae

H Rosaceae

H Rutaceae

H Scrophulariaceae
Sopubia trifida Buch. Ham ex G.Don

H Verbenanaceae
Lantana camara L.
Lp LILIOPSIDA H Phormiaceae
33 34 35 36 37 38

H Poaceae
Hng bi Trc tho lng Xoang trc sc Tc hnh tm San trn Sy nam
68
Chamae. Chamae. Chamae. Chamae. Chamae. Chamae.
Arudinella setosa Trin. Coelorachis striata (Steud.) A. Cam. Digitaria violascens Link Paspalum orbiculare Forst. Phragmites australis (Cav.) Trin. ex steud.
39
Rhynchelytrum repens (Willd.) C.E. Hubb.
Hng nhung
Thero.
DL01
V tr: 110 2320,6 v Bc 1080 0522,0 kinh ng cao so vi mt nc bin : 890m dc Hng dc S loi (S) S c th (N) - Cy g ln : 15 - Cy g nh : 95 - Cy dng bi : 852 - Cy dng c : 25.725 Tng s c th ca DL01, N = 26.687 DMarg = 3,92 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DL01
: 250 : -T : 41 loi
Tn vit loi
Dng sng
Tnh trng
1 2 3
Adiantum flabellatum L.
H Blechnaeae
Cy vt, rn en Rng da ng Seo g, chn x
Cryp. Nano. Cryp.
Blechnum orientale L.
H Pteridaceae
Pteris vittata L.

Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Asclepiadaceae
69
Thng ba l
Macro.
5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25

H Asteraceae
H th trng Sn cc nhm B ch, bp l Tai ngh rng Sc daltonii Bic l to Trng qu bi ng ba tri c cng u sam Mc c
Lian. Chamae. Nano. Micro. Micro. Lian. Thero. Lian. Lian. Chamae. Micro. Micro. Chamae. Nano. Nano. Chamae. Nano. Nano. Nano. Nano. Nano.
Wedenia urticaefolia (Blume) DC. ex Wight

H Buddleiaceae

H Euphorbiaceae

H Fabaceae
Clitoria macrophylla Wall. ex Benth. Desmodium adscendens (Sw.) DC. Dunbaria polocarpa Kurz. Dunbaria villosa (Thunb.) Makino. Mimosa pudica L.
H Fagaceae

H Juglandaceae
D trng Cho bng Gi hc Mua lng Sm nhn An bch Trung Quc Cm ngui rng Hng sim Mm xi Ngy hng Dt snh gn

H Leeaceae

H Melastomataceae
Melastoma saigonense (Kuntze) Merr. Memecylon acuminatun Smith ex Triana Osbeckia chinenensis L.
H Myrsinaceae

H Myrtaceae
Rhodomyrtus tomentosa (Ait.) Hassk.

H Rosaceae

H Rubiaceae
Pavetta nervosa Craib.
70
H Theaceae
26 27

H Tiliaceae
Giang ni C ke lng
Micro. Nano.
Grewia hirsuta Vahl.

28 29 30 31 32 33 34 35 36 37 38 39 40 41
H Cyperaceae
Trai thng Mao th st Cng Kerr Hng bi Song on Ct v p L tho gn C tranh K ni L tam hng t, cht C ui nhn Th phc linh Gng
Thero. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Cryp. Thero. Chamae. Chamae. Thero. Lian. Cryp.
Fimbristylis ferruginea (L.) Vahl. Scleria kerrii Turrill

H Phormiaceae

H Poaceae
Diectomis fastigiata (Sw.) Kunth Eulalia speciosa (Deb.) Kuntze Ichnanthus vicinus (F.M.Bail.) Merr. Imperata cylindrica (L.) P.Beauv. Panicum notatum Retz. Themeda triandra Forssk. Thysanolaena maxima (Roxb.) Kuntze Urochloa paspaloides J & C, Presl
H Smilacaceae

H Zingiberaceae
DL02
V tr: 110 2644,8 v Bc 1080 0342,1 kinh ng cao so vi mt nc bin : 1.172m dc Hng dc : 500 : N-TB
71
S loi (S) S c th (N) - Cy g ln : 25 - Cy g nh : 58 - Cy dng bi : 752
: 42 loi
- Cy dng c : 23.275 Tng s c th ca DL02, N = 24.110 DMarg = 4,06 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DL02
Tn vit loi
Dng sng
Tnh trng
1 2 3 4 5

H Aspleniaceae
Rng nguyt s Rng can x hnh gm Rng i dc Gut, t, rng ty sn ui phng l si
Cryp. Epi. Cryp. Cryp. Epi.
Asplenium ensiforme Wall. Ex HooK. & Grev.

H Dennstaedtiaceae

H Gleicheniaceae

H Polypodiaceae
Drynaria quercifolia (L.) J. Smith


Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Asclepiadaceae
Thng ba l
Macro.
7 8 9

H Asteraceae
H th trng Chua l nhm Bch u
Lian. Thero. Chamae.
Emilia scabra DC. Vernonia squarrosa (D.Don) Less.

H Campanulaceae
72
10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31

H Dilleniaceae
ng sm Java Dy chiu Cp mc hnh sao Ca di xoan Tai ngh rng Sc daltonii Me rng B ngt hai mu Doi Chevalier Bic l to Trng qu Griffith. Chm en u ba thy D trng Mua lng Sm nhn Cm ngui l n mng Dum Trung b Tr hu An tc Chn tr Harman
Lian. Lian. Micro. Micro. Micro. Micro. Micro. Nano. Micro. Lian. Chamae. Nano. Lian. Micro. Nano. Nano. Nano. Nano. Nano. Micro. Micro. Nano.

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Lyonica ovalifolia (Wall.) Drude

H Euphorbiaceae
Aporusa serrate Gagnep. Glochidion daltonii (Muel-Arg.) Kurz. Phyllanthus em blica L Sauropus bicolor Craib
H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Clitoria macrophylla Wall. ex Benth. Desmodium griffithianum Benth. Indigofera nigrescens Kurz. ex King & Prain Vigna triloba (L.) Ver dc.
H Fagaceae

H Melastomataceae
Melastoma saigonense (Kuntze) Merr. Memecylon acuminatun Smith ex Triana

H Myrsinaceae
Ardisia mirabilis Pit. Maesa membranacea A.DC.

H Rosaceae

H Rubiaceae
Wendlantia glabrata DC.

H Styracaceae

H Theaceae

Lp LILIOPSIDA
73
H Arecaceae
32
Phoenix loureiri Kunth. Var. humilis (Becc.) S.C.Barow.

H Cyperaceae
Ch l nh
Chamae
33 34 35 36 37 38 39 40 41 42
Carex lindleyana Nees

H Phormiaceae
Kit Hng bi C may vng Xoang trc ng c Mao phng chm C tranh Mm ru C phao Th phc linh Gng
Chamae. Chamae. Chamae. Chamae. Chamae. Cryp. Chamae. Chamae. Lian. Cryp.

H Poaceae
Chrysopogon fulvus (Spreng.) Chiov. Coelorachis pratensis (Bal.) A.Cam. Erianthus fastigiatus Nees ex Steud Imperata cylindrica (L.) P.Beauv. Ischaemum barbatum Retz. Themeda caudata (Nees.) HacK.
H Smilacaceae

H Zingiberaceae

DL03
V tr: 110 2537,1 v Bc 1080 0332,9 kinh ng cao so vi mt nc bin : 1.032m dc Hng dc S loi (S) S c th (N) - Cy g ln : 32 - Cy g nh : 121 - Cy dng bi : 952 - Cy dng c : 24.500 Tng s c th ca DL03, N = 25.605 DMarg = 4,23 : 50 : -T : 44 loi
74
Bng: Danh lc thnh phn loi, dng sng v tnh trng loi DL03
Tn vit loi
Dng sng
Tnh trng
1 2 3 4
Adiantum flabellatum L. Taenitis blechnoides (Willd.) SW.

H Polypodiaceae
Cy vt, rn en Rng i dc rng Chn x Hy Lp
Cryp. Cryp. Epi. Cryp.
Aglaomorpha coronans (Wall. Ex Mett.) Copel.

H Pteridaceae
Pteris cretica L.

Thng ba l
Macro.
6 7 8 9 10 11 12 13 14 15 16 17
Eupatorium odoratum L.
H Ericaeae
C Lo, c cng sn Cp mc hnh sao Nem l liu Chi mi Walker Tai ngh rng Sc daltonii B ngt hai mu Doi Chevalier Gip qu u l nh
Thero. Micro. Micro. Nano. Micro. Micro. Nano. Micro. Lian. Thero. Micro. Micro.

H Euphorbiaceae
Antidesma walkerii Pax & Hoffm. Aporusa serrate Gagnep. Glochidion daltonii (Muel-Arg.) Kurz. Sauropus bicolor Craib
H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Cajanus elongatus (Benth) Maesen Desmodium sp.
H Fagaceae
Lithocarpus parvulus (Hickel & A.Camus) A.Camus

H Juglandaceae
D gi Cho bng
75
H Lamiaceae
18 19 20
Nosema cochinchinensis (Lour.) Merr.

H Melastomataceae
Cm thy trung vit Mua trng Du ru
Thero. Nano. Micro.
Melastoma candidum D.Don

H Myricaceae
H Myrsinaceae
21 22 23 24 25 26 27 28 29 30 31
Ardisia annamensis Pit. Ardisia crenta Sims.

H Myrtaceae
Cm ngui Trung b Cm ngui rng Hng sim Dum Trung b Dt snh gn Gii lm, du da An tc Chn tr Harman T chu T th c ui Hoa tm n
Nano. Nano. Nano. Nano. Nano. Nano. Micro. Nano. Nano. Lian. Chamae.
Rhodomyrtus tomentosa (Ait.) Hassk.

H Rosaceae

H Rubiaceae

H Rutaceae

H Styracaceae

H Theaceae

H Verbenanaceae

H Vitaceae
Tetrastigma caudatum Merr. & Chun.

H Violaceae
Viola inconspicua Blume

Lp LILIOPSIDA H Arecaceae
32 33 34
Caryota sympelata Gagn. Murdannia giganteum ( Vahl.) Bruckner

H Cyperaceae
ng nh Trai cao Bm dy
Chamae. Thero. Thero.
Bulbostylis densa (Wall.) Hand. Mazz.
76
35 36 37 38 39 40 41 42 43 44
Carex lindleyana Nees Fimbristylis pubisquama Kern

H Phormiaceae
Kit Mao th vy Hng bi Tc hnh ht Tinh tho st Ct v ra K Hayata C bng lau L tam hng Kim cang lin hng
Chamae. Chamae. Chamae. Thero. Chamae. Chamae. Chamae. Chamae. Chamae. Lian.
Dianella ensifolia (L.) DC. Digitaria abludens (Roem. & Sch.) Veldk Eragrostis ferruginea (Thumb.) P.Beauv. Eulalia fimbriata (Hack.) Kuntze Panicum hayatae A.Cam Saccharum spontaneum L. Themeda triandra Forssk.
H Smilacaceae

NM01
V tr: 110 4948,9 v Bc 1080 3843,9 kinh ng cao so vi mt nc bin : 901m dc Hng dc S loi (S) S c th (N) - Cy g ln : 16 - Cy g nh : 48 - Cy dng bi : 680 - Cy dng c : 39.200 Tng s c th ca NM01, N = 39.944 DMarg = 3,68 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi NM01
: 300 : TN-B : 40 loi
Tn vit loi
Dng sng
Tnh trng
77
Cy vt, rn en
Cryp.
H Blechnaeae

Rng bit x
Nano.

Thng ba l
Macro.
4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20
Ageratum conyzoides L.
H Asteraceae
C ct heo Chua l nhm Rau tu bay Sn cc ni B ch, bp l Dy chiu Tai ngh rng D fleury Me rng Trng qu tc vng Thc lp Schubert Mc c
Chamae. Thero. Thero. Thero. Nano. Lian. Micro. Nano. Micro. Thero. Nano. Chamae. Micro. Nano. Nano. Nano. Micro.
Emilia scabra DC. Gynura crepidoides Benth. Wedenia wallichii Lees

H Buddleiaceae

H Dilleniaceae

H Euphorbiaceae
Aporusa serrate Gagnep. Breynia fleuryi Beille Phyllanthus em blica L

H Fabaceae
Desmodium auricomum Grah. ex Benth. Desmodium schubertiae Ohashi Mimosa pudica L.

H Fagaceae

H Lamiaceae
D trng Cha d Mua lng Sm nhn Du ru

H Melastomataceae
Melastoma saigonense (Kuntze) Merr. Memecylon acuminatun Smith ex Triana

H Myricaceae

78

H Myrsinaceae
21 22 23 24 25 26 27 28 29 30 31 32
Ardisia crenta Sims. Maesa membranacea A.DC.

H Proteaceae
Cm ngui rng n mng Qun hoa cao Ngy hng Dt snh gn Gii lm, du da ni Ava Chn tr Harman Giang ni C ke lng T chu T th c ui
Nano. Nano. Micro. Nano. Nano. Nano. Thero. Nano. Micro. Nano. Nano. Lian.

H Rosaceae

H Rubiaceae

H Rutaceae

H Scrophulariaceae

H Theaceae
Eurya Japonica var. harmandii Pierre ex Pitard Ternstroemia japonica (Thunb.) Thunb.
H Tiliaceae

H Verbenanaceae

H Vitaceae

33 34 35 36 37 38
Caryota sympelata Gagn.

H Cyperaceae
ng nh Mao th trn di Hng bi Mao cc hoa nh Ct v p C tranh

79
Chamae. Thero. Chamae. Chamae. Chamae. Cryp.

H Phormiaceae

H Poaceae
Capillipedium parviflorum (R.Dr.) Stapf Eulalia speciosa (Deb.) Kuntze Imperata cylindrica (L.) P.Beauv.
39 40
Rhynchelytrum repens (Willd.) C.E. Hubb. Seraria parviflora (Poir.) Roem. & Schult.
NM02
Hng nhung ui chn
Thero. Chamae.
V tr: 110 4956,7 v Bc 1080 3840,5 kinh ng cao so vi mt nc bin : 960m dc Hng dc S loi (S) S c th (N) - Cy g ln : 17 - Cy g nh : 65 - Cy dng bi : 750 - Cy dng c : 44.100 Tng s c th ca NM02, N = 44.932 DMarg = 3,73 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi NM02
: 150 : B-N : 41 loi
Tn vit loi
Dng sng
Tnh trng
1 2 3
H Blechnaeae
Cy vt, rn en Rng da ng Rng tai chut li dao
Cryp. Nano. Epi.
Blechnum orientale L.
H Polypodiaceae


Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Anacardiaceae
Thng ba l
Macro.
80
5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23

H Apriaceae
Mui, Ng bi t Rau m H th trng Chua l nhm Rau tu bay Dc cnh cnh Sn cc ni Dy chiu Cp mc hnh sao Tai ngh rng Me rng Sc sc cao Trng qu tc vng D trng Si dui Cha d Gi hc Sm nhn Du ru
Micro. Chamae. Lian. Thero. Thero. Thero. Thero. Lian. Micro. Micro. Micro. Nano. Thero. Micro. Micro. Nano. Chamae. Nano. Micro.

H Asclepiadaceae

H Asteraceae
Emilia scabra DC. Gynura crepidoides Benth. Laggera alata (D.Don) Schultz Bip ex Oliv. Wedenia wallichii Lees
H Dilleniaceae

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm

H Euphorbiaceae
Aporusa serrate Gagnep. Phyllanthus em blica L

H Fabaceae
Crotalaria angyroides H.B.K Desmodium auricomum Grah. ex Benth.

H Fagaceae
Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus setulosa Hickel& A. Camus
H Lamiaceae

H Leeaceae

H Melastomataceae
Memecylon acuminatun Smith ex Triana

H Myricaceae
H Myrsinaceae
24

81
Cm ngui rng
Nano.
H Rosaceae
25 26 27 28 29 30 31 32

H Rubiaceae
Dum Trung b Dt snh gn Ba chc ni Ava Con chut Giang ni C roi nga T th c ui
Nano. Nano. Nano. Thero. Nano. Micro. Phane. Lian.

H Rutaceae

H Scrophulariaceae

H Sterculiaceae
Helicteres hirsuta Lour.

H Theaceae

H Verbenanaceae
Verbena officinalis L.
H Vitaceae

33
Phoenix loureiri Kunth. Var. humilis (Becc.) S.C.Barow.

H Cyperaceae
Ch l nh
Chamae.
34 35 36 37 38 39 40 41
Fimbristylis oblonga T.Koyama Scleria kerrii Turrill

H Poaceae
Mao th trn di Cng Kerr Mao cc hoa nh Gt nm C tranh Hng nhung ui chn t, cht
Thero. Chamae. Chamae. Chamae. Cryp. Thero. Chamae. Chamae.
Capillipedium parviflorum (R.Dr.) Stapf Garnotia barbulata (nees) Merr. Imperata cylindrica (L.) P.Beauv. Rhynchelytrum repens (Willd.) C.E. Hubb. Seraria parviflora (Poir.) Roem. & Schult. Thysanolaena maxima (Roxb.) Kuntze
NM03
V tr: 110 5010,6 v Bc 1080 3913,1 kinh ng

82
cao so vi mt nc bin : 815m dc Hng dc S loi (S) S c th (N) - Cy g ln : 16 - Cy g nh : 74 - Cy dng bi : 682 - Cy dng c : 35.525 Tng s c th ca NM03, N = 36.297 DMarg = 3,62 Bng: Danh lc thnh phn loi, dng sng v tnh trng loi NM03
: 350 : -T : 39 loi
Tn vit loi
Dng sng
Tnh trng
1 2
Taenitis blechnoides (Willd.) SW.

H Blechnaeae
Rng i dc Rng bit x
Cryp. Nano.


Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Anacardiaceae
Thng ba l
Macro.
4 5 6 7 8 9

H Asclepiadaceae
Mui, Ng bi t H th trng Chua l nhm Rau tu bay Bu t, tam tht gi Sn cc ni

83
Micro. Lian. Thero. Thero. Chamae. Thero.

H Asteraceae
Emilia scabra DC. Gynura crepidoides Benth. Gynura divaricata (L.) DC. Wedenia wallichii Lees
H Buddleiaceae
10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26

H Ericaeae
B ch, bp l Ca di xoan Tai ngh rng Sc daltonii Ba bt trng Me rng Sc sc cao ng ba ngn D trng Cau phong lun Gi hc Mua trng Cm ngui rng Dum Trung b Tr hu Gii lm, du da nhiu gai
Nano. Micro. Micro. Micro. Nano. Micro. Nano. Lian. Micro. Thero. Chamae. Nano. Nano. Nano. Micro. Nano.
Lyonica ovalifolia (Wall.) Drude

H Euphorbiaceae
Aporusa serrate Gagnep. Glochidion daltonii (Muel-Arg.) Kurz. Mallotus apelta (Lour.) Muell. Arg. Phyllanthus em blica L
H Fabaceae
Crotalaria angyroides H.B.K Dunbaria fusca (Wall.) Kurz.

H Fagaceae

H Lamiaceae
Clinopodium gracile (Benth.) Matsum.

H Leeaceae

H Melastomataceae
Melastoma candidum D.Don

H Myrsinaceae

H Rosaceae

H Rubiaceae
Wendlantia glabrata DC.

H Rutaceae
Clausena excavata Burm.f. Zanthoxylum myriacanthum Wall. ex HooK.f.

H Sterculiaceae
Cm ru Hong mc Nano.
27 28
H Theaceae
kn Chn tr Harman
Nano. Nano.

84
H Tiliaceae
29 30

H Verbenanaceae
C ke lng C roi nga
Nano. Phane.
Verbena officinalis L.
31 32 33 34 35 36 37 38 39
Caryota sympelata Gagn.

H Commelinaceae
ng nh Trai thng Lc ta Trc tho lng Tinh tho st C tranh K Hayata Hng nhung Kim cang lin hng
Chamae. Thero. Thero. Chamae. Chamae. Cryp. Chamae. Thero. Lian.
H Cyperaceae
Cyperus digitatus Roxb.

H Poaceae
Arudinella setosa Trin. Eragrostis ferruginea (Thumb.) P.Beauv. Imperata cylindrica (L.) P.Beauv. Panicum hayatae A.Cam Rhynchelytrum repens (Willd.) C.E. Hubb.
H Smilacaceae
III. KT QU NGHIN CU TNG HP V A DNG SINH HC THC VT KIU RNG THNG BA L (Pinus kesiya) MC T NHIN LM NG V CNG LN CN. Bng cc ch s v a dng thc vt 20 tiu chun ca kiu rng thng ba l (Pinuskesiya) mc t nhin tnh Lm ng v vng ln cn
cao so K hiu V Kinh vi mt nc bin (m) LB01 LB02 LB03 TL01 TL02 120 0223,7 120 0227,1 120 0228,8 110 530,2 110 5252,2 1080 2538,1 1080 2532,9 1080 2540,8 1080 2557,0 1080 2558,3 1794 1988 1890 1462 1553
85
dc, Hng dc 300, TB-N 300, T- 120, TB-N 150, N-B 320, TN-B
S loi (S) 40 39 41 42 39
S c th (N) 43.728 44.880 39.024 40.163 34.993 Ch s DMarg.
3,65 3,55 3,78 3,87 3,63
TL03 NV01
110 5242,3 110 5221,4
1080 2610,2 1080 2616,3 1080 2728,9 1080 2729,3 1080 2231,6 1080 2150,4 1080 2202,0 1080 2912,5 1080 2917,4 1080 0522,0 1080 0342,1 1080 0332,9 1080 3843,9 1080 3840,5 1080 3913,1
1383 1334 1228 1310 1378 1427 1455 1459 1517 890 1172 1032 901 960 815
100, TN-B 150, N-B 280, N-B 350, N-TB 150, TN-B 350, N-B 250, N-TB 400, ,N-TTB 250, TN-B 250, -T 50, N-TB 50, -T 300, TB-N 150, B-N 350, -T
41 43 42 39 40 43 41 42 39 41 42 44 40 41 39
51.197 52.515 48.605 39.977 53.626 43.882 48.751 49.195 36.084 26.687 24.110 25.605 39.944 44.932 36.297
3,69 3,86 3,80 3,59 3,58 3,93 3,71 3,82 3,62 3,92 4,06 4,23 3,68 3,73 3,62
DTL01 110 5425,5 DTL02 110 5417,4 TN01 SV01 SV02 DS01 DS02 DL01 DL02 DL03 NM01 NM02 NM03 110 5559,8 110 5918,5 110 5914,6 120 0046,8 120 0045,7 110 2320,6 110 2644,8 110 2537,1 110 4948,9 110 4956,7 110 5010,6
Cc ch s nu trn l kt qu nghin cu 20 tiu chun c thit lp trong cc khi rng thng ba l cao 800m n 2.000m. Qua bng trn ta thy ch s a dng Margalef ca cc chnh lch nhau khng nhiu, to thnh tp hp s kh ng nht. V vy ch s trung bnh cng c th i din mt cch y v cht ch cho tp hp cc . Ch s Margalef chung cho kiu rng thng ba l mc t nhin Lm ng v vng ln cn l: DMarg = 3,76. ng thi, nh phn phng php nghin cu t ra l cc phi cch xa mp rng, v rng thng y phi c cu trc 3 tng, nhng rng thng ny ang trong trng thi pht trin t nhin nht. C th ni rng nhng cnh rng ny ang giai on Climax ca din th, chng c thnh phn v dng sng ca cc loi l kh n nh. nhiu c th ca cc loi cng kh n nh v c nh gi trn c s thang ca Drude. T cc kt qu nghin cu cc , chng ti tng hp thnh danh lc thc vt cho kiu rng thng ba l:
86
BNG DANH LC THNH PHN LOI, DNG SNG, TNH TRNG V NHIU CA LOI KIU RNG THNG BA L MC T NHIN LM NG V VNG LN CN. TT NGNH, H TN KHOA HC LOI Ngnh LYCOPDIOPHYTA H Lycopodiaceae TN VIT LOI DNG SNG TNH TRNG NHIU
1 2
Lycopodionella cernuua (L.) Pic.Serm.

H Selaginellaceae
Thch tng nghin Quyn b n bo t
Thero. Thero.
Sol Sol

Ngnh POLYPODIOPHYTA H Adiantaceae
3 4 5 6 7
Adiantum flabellatum L. Adiantum stenochlamys Bak. Taenitis blechnoides (Willd.) SW.

H Aspleniaceae
Cy vt, rn en Rng nguyt s C nc Rng can x gn Rng can x hnh gm
Cryp. Cryp. Cryp. Epi. Epi.
Cop 1 Cop 1 Sp Sp Sol
Asplenium affine Sw. Asplenium ensiforme Wall. Ex HooK. & Grev.

H Blechnaeae
8 9 10
Blechnum orientale L. Brainea insignis ( HooK.) J. Smith Woodwardia Japonica (L.f.) J.Sm.
H Dennstaedtiaceae
Rng da ng Rng bit x Rng bch hoa nht
Nano. Nano. Chamae
Sol Cop 1 Cop 1
11 12
Microlepia platyphylla (D.Don) J.Smith Pteridium aquilinum (L.) Kuhn.

H Dipteridaceae
Rng vi ln Rng i dc
Cryp. Cryp.
Sol Cop 3
13 14 15
Dipteris conjugata (Kaulf.) Reinw.

H Gleicheniaceae
Song dc i Gut, t, rng ty sn Rng t minh qu
Cryp. Cryp. Cryp.
Sol Cop 3 Sol

H Osmundaceae
Osmunda cinnamomea L.
87
H Polypodiaceae
16 17 18 19 20 21 22
Aglaomorpha coronans (Wall. Ex Mett.) Copel. Crypsinus rhynchoplyllus (HooK.) Copel. Drynaria quercifolia (L.) J. Smith Gonophlebium subauriculatum (Bl.) Prest. Lemmaplyllum carnosum (HooK.) C. Presl. Paragramma banaensis (C. Chr). Ching Pyrrosia lingua (Thunb.) Farw. Var. lingua
H Pteridaceae
rng Rng n thy c mi ui phng l si Rng a tc Rng vy c nc Rng song vn B N Rng tai chut li dao
Epi. Epi. Epi. Epi. Epi. Epi. Epi.
Sol Sol Sol Sol Sol Sp Cop 1
23 24
Pteris cretica L. Pteris vittata L.

Chn x Hy Lp Seo g, chn x
Cryp. Cryp.
Sol Sol
25 26
Keteleeria evelyniana Mast Pinus kesiya Royle ex Gordon

Ngnh MAGNOLIOPHYTA Lp Magnoliopsida H Acanthaceae
Du sam ni t Thng ba l
Macro. Macro.
Sp Soc
27 28 29 30 31 32 33
Lepidagathis hyalina Ness

H Anacardiaceae
Ln chy Mui, Ng bi t Rau m Rau m chevalier Bng bin H da l to H th trng
Thero. Micro. Chamae. Chamae Chamae Epi. Lian.
Sol Cop 1 Sp Sol Sol Sol Cop 1

H Apriaceae
Centella asiatica (L.)Urb. Hydrocotyle chevalieri (Chern) Tard. Pimpenella diversifolia DC.
H Asclepiadaceae
Hoya macrophylla Blume. Streptocaulon juventas (Lour.) Merr.

H Asteraceae
34 35
Ageratum conyzoides L. Ainsliaea petelotii Merr.
C ct heo Anh l petelot.
Chamae. Chamae.
Cop 1 Sol
88
36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61
Anaphalis lactea Maxim Bidens pilosa L. Blumea sinuata (Lour.) Merr. Colobogyne langbianense Gagn. Conyza canadense (L.) Crong. Dichrocephala integrifolia (L.f.) Kuntre Elephantopus mollis H.B.K Elephantopus scaber L. Emilia scabra DC. Emilia sonchifolia (L.) DC Eupatorium odoratum L. Galingsoga parviflora Cav. Gerbera piloselloides (L.) Cass. Gnaphalium luteo-album L. Gynura crepidoides Benth. Gynura divaricata (L.) DC. Hypochaeris radicata L. Inula nervosa Wall.ex DC. Laggera alata (D.Don) Schultz Bip ex Oliv. Sonchus oleraceus L. Spilanthes paniculata wall. ex DC. Vernonia squarrosa (D.Don) Less. Wedenia urticaefolia (Blume) DC. ex Wight Wedenia wallichii Lees
H Berberidaceae
Bch nhung sa n but, i bi l ln Riu Thng lo, tai hm Lng st l nguyn Chn voi mm, cao Chn voi nhm Chua l nhm C chua l C Lo, c cng sn Vi cc Cc lng, Khc vng Rau tu bay Bu t, tam tht gi Miu nh Inugn Dc cnh cnh Nh cc Cc o, nt o Bch u Sn cc nhm Sn cc ni Hong lin r Khuyn thit
Thero. Thero. Thero. Thero. Thero. Thero. Chamae. Chamae. Thero. Thero. Thero. Thero. Chamae. Thero. Thero. Chamae. Chamae. Thero. Thero. Thero. Thero. Chamae. Chamae. Thero. Nano. Thero.
Sol Cop 1 Sol Cop 1 Cop 1 Sol Sol Cop 1 Cop 1 Sol Cop 1 Sp Sol Cop 1 Cop 2 Sol Sol Sol Cop 1 Sol Sol Cop 1 Cop 1 Cop 2 Sol Sol
Mahonia nepalensis DC.

H Boraginaceae
Cynoglossum zeylanicum (Vahl. ex Hornem.) Thunb. ex Lehm.

H Buddleiaceae
62
B ch, bp l
Nano.
Cop 2
89
H Campanulaceae
63 64 65 66 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 86 87

H Caprifoliaceae
ng sm Java Vt dai Si nht Dy chiu Bt rui Trng l bn nguyt Cp mc hnh sao Ca di xoan Nem l liu Chi mi Walker Tai ngh rng D fleury Sc daltonii Ba bt trng Dip h chu trng Me rng B ngt hai mu Doi Chevalier Gip qu Bin hng rng thng Mung trinh n Bic l to Sc sc trng Sc sc cao Trng qu bi
90
Lian. Micro. Thero. Lian. Thero. Thero. Micro. Micro. Micro. Nano. Micro. Nano. Micro. Nano. Thero. Micro. Nano. Micro. Lian. Nano. Thero. Lian. Chamae. Nano. Thero.
Cop 1 Sp Cop 2 Cop 1 Sp Sp Cop 3 Cop 2 Cop 2 Sp Cop 3 Sp Cop 3 Sp Sol Cop 3 Cop 1 Cop 3 Sp Cop 1 Cop 1 Cop 1 Cop 1 Sp Sp

H Chloranthaceae

H Dilleniaceae

H Draseraceae
Drosera burmannii Vahl. Drosera peltata Smith in Willd.

H Ericaeae
Craibiodendron stellatum (Pierre ex Lanees) W.W.Sm Lyonica ovalifolia (Wall.) Drude Vaccinium iteophyllum Hance
H Euphorbiaceae
Antidesma walkerii Pax & Hoffm. Aporusa serrate Gagnep. Breynia fleuryi Beille Glochidion daltonii (Muel-Arg.) Kurz. Mallotus apelta (Lour.) Muell. Arg. Phyllanthus amarus Schum. & Thonn. Phyllanthus em blica L Sauropus bicolor Craib
H Fabaceae
Archidendron chevalieri (Kost.) I. Niels. Cajanus elongatus (Benth) Maesen Campylotropis pinetorum (Kurz) Schindl. Cassia mimosoides L. Clitoria macrophylla Wall. ex Benth. Crotalaria albida Heyne ex Roth. Crotalaria angyroides H.B.K Desmodium adscendens (Sw.) DC.
88 89 90 91 92 93 94 95 96 97 98 99
Desmodium auricomum Grah. ex Benth. Desmodium concinnum DC. Desmodium griffithianum Benth. Desmodium multiflorum DC. Desmodium schubertiae Ohashi Desmodium sp. Desmodium umbellatum (L.) Dc. Dunbaria fusca (Wall.) Kurz. Dunbaria polocarpa Kurz. Dunbaria villosa (Thunb.) Makino. Flemingia lincata var. glutinosa Prain Flemingia macophylla (Willd.) Prain
Trng qu tc vng Trng qu ngh thut Trng qu Griffith. Trng qu nhiu hoa Thc lp Schubert u l nh Trng qu tn ng ba ngn ng ba tri c cng u sam Tp m l nh Tp m l to Chm ui di Chm en Chm cua Mc c u ma Mang sang on kim ta Hu v hoe u ba thy D trng D gi Si Helfer. Si kerr Si lng Si dui Long m Long m Langbian
91
Thero. Nano. Chamae. Chamae. Nano. Thero. Thero. Lian. Lian. Lian. Nano. Nano. Nano. Nano. Nano. Chamae. Lian. Thero. Chamae. Chamae. Lian. Micro. Micro. Micro. Micro. Micro. Micro. Thero. Thero.
Cop 1 Sp Cop Cop 1 Cop 1 Sol Cop 2 Sp Cop 2 Cop 1 Cop 2 Cop 1 Sp Cop 1 Cop 2 Cop 1 Sp Sp Cop 1 Sp Sp Cop 3 Sol Cop 2 Cop 1 Cop 3 Cop 1 Cop 1 Sol
100 Indigofera longicandata Thun 101 Indigofera nigrescens Kurz. ex King & Prain 102 Indigofera stachyodes Lindl. 103 Mimosa pudica L. 104 Pueraria phaseoloides (Roxb.) Benth. 105 Shuteria suffutta Benth. 106 Tephrosia purpurea (L.) Pers. 107 Uraria rufescens (DC.) Schindl. 108 Vigna triloba (L.) Ver dc.
H Fagaceae
109 Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. 110 Lithocarpus parvulus (Hickel & A.Camus)
A.Camus
111 Quercus helferiana A.DC. 112 113 114 Quercus kerri Craib. Quercus lanata Smith in Rees. Quercus setulosa Hickel& A. Camus
H Gentianaceae
115 Gentiana indica Steud. 116 Gentiana langbianensis A, Chev. ex H.
Smith
H Juglandaceae
117 Engelhardia spicata Lesch. ex Blume

H Lamiaceae
Cho bng Cau phong lun Cha d Hng nhu x Bch thit Cm thy trung vit Lin n gi Gi hc Sc mo ct Lun tho l trn
Micro. Thero. Nano. Nano. Thero. Thero. Micro. Chamae. Thero. Thero.
Cop 1 Sp Cop 1 Cop 1 Cop 1 Sp Cop 1 Cop 2 Sol Sp
118 Clinopodium gracile (Benth.) Matsum. 119 Elsholtzia blanda (Benth.) Benth. 120 Elsholtzia winitiana Craib 121 Leucas ciliata Benth. 122 Nosema cochinchinensis (Lour.) Merr.
H Lauraceae
123 Lindera spicata Kosterm.

H Leeaceae
124 Leea rubra Bl. ex Spreng.

H Loganiaceae
125 Mitrasaeme eriophila Leenh.

H Lythraceae
126 Rotala rotundifolia (HooK.f. ex Roxb) Koehne

H Melastomataceae
127 Melastoma candidum D.Don 128 Melastoma chevalieri Guill. 129 Melastoma saigonense (Kuntze) Merr. 130 Memecylon acuminatun Smith ex Triana 131 Osbeckia chinenensis L. 132 Osbeckia cupulasis D.Don ex W. & Arn. 133 Osbeckia nepalensis HooK. 134 Osbeckia stellata Buch. Ham. ex D.Don H Moraceae 135 Ficus hirta Vahl var. roxburghii (Miq.) King
H Myricaceae
Mua trng Mua chevalier Mua lng Sm nhn An bch Trung Quc An bch u An bch Nepal An bch sao Ngi kh
Nano. Nano. Nano. Nano. Chamae. Thero. Thero. Chamae. Nano.
Cop 3 Sp Cop 3 Cop 1 Cop 2 Cop 1 Cop 1 Cop 1 Sp
136 yrica esculenta Buch. Ham. ex D.Don var. Chevalieri (Dode) Pham Hoang
Du ru
Micro.
Cop 3
92
H Myrsinaceae
137 Ardisia annamensis Pit. 138 Ardisia crenta Sims. 139 Ardisia mirabilis Pit. 140 Maesa membranacea A.DC.
H Myrtaceae
Cm ngui Trung b Cm ngui rng Cm ngui l n mng
Nano. Nano. Nano. Nano.
Cop 2 Cop 2 Cop 1 Cop 1
141 Rhodomyrtus tomentosa (Ait.) Hassk.

H Nepenthacece
Hng sim Bnh nc Trung b Me t nh M Kch nh, du nng Qun hoa cao Mm xi Dum Trung b Ngy hng
Nano. Chamae. Chamae. Chamae. Thero. Micro. Nano. Nano. Nano.
Sp Cop 1 Cop 1 Sol Cop 1 Cop 3 Cop 2 Cop 3 Cop 2
142 Nepenthes annamensis Macfarl.

H Oxalidaceae
143 Oxalis corniculata L

H Plantaginaceae
144 Plantago asiatica L.

H Polygalaceae
145 Polygala paniculata L.

H Proteaceae
146 Helicia excelsa (Roxb.) Blume

H Rosaceae
147 Rubus alceaefolius Poir. 148 Rubus annamensis Card. 149 Rubus cochinchinensis Traht.
H Rubiaceae
150 Hedyotis auricularia L. 151 Hedyotis rudis (Pierre ex Pit.) Phamh. 152 Pavetta nervosa Craib. 153 Wendlantia glabrata DC.
H Rutaceae
An in tai An in nhm Dt snh gn Tr hu
Thero. Thero. Nano. Micro.
Cop 1 Cop 1 Cop 1 Cop 1
154 Clausena excavata Burm.f. 155 Euodia lepta (Spreng.) Merr. 156 Glycosmis pentaphylla (Retz.) DC. 157 Zanthoxylum myriacanthum Wall. ex HooK.f.
H Scrophulariaceae
Gii lm, du da xoan Ba chc Cm ru Hong mc nhiu gai
Nano. Nana. Nano. Nano.
Cop 2 Cop 1 Cop 1 Sol
93
158 Alectra arvenses (Benth.) Merr. 159 Sopubia trifida Buch. Ham ex G.Don
H Sterculiaceae
ni Ava S bu ch ba kn Con chut An tc Chn tr Harman Giang ni
Thero. Thero. Nano. Nano. Micro. Nano. Micro.
Cop 2 Sp Cop 1 Sp Cop 1 Cop 2 Cop 3
160 Helicteres angustifolia L. 161 Helicteres hirsuta Lour.

H Styracaceae
162 Styrax benjoin Dryand

H Theaceae
163
164 Ternstroemia japonica (Thunb.) Thunb.

H Tiliaceae
165 Grewia hirsuta Vahl. 166 Triumfetta pseudocana Sprague & Craib.
H Verbenanaceae
C ke lng Gai u lng
Nano. Nano.
Cop 1 Sol
167 Callicarpa rubella Lindl. 168 Lantana camara L. 169 Verbena officinalis L.
H Vitaceae
T chu Ng sc C roi nga
Nano. Nano. Phane.
Cop 1 Sp Sp
170 Tetrastigma caudatum Merr. & Chun.

H Violaceae
T th c ui Hoa tm n
Lian. Chamae.
Cop 1 Sp
171 Viola inconspicua Blume

Lp LILIOPSIDA H Amaryllidaceae
172 Hypoxis aurea Lour.

H Arecaceae
H trm, tin mao ng nh Ch l nh
Chamae. Chamae. Chamae.
Sol Sp Sp
173 Caryota sympelata Gagn. 174 Phoenix loureiri Kunth. Var. humilis (Becc.) S.C.Barow.
H Asparagaceae
175 Asparagus filicinus Buch Ham ex D.Don. Thin mn rng

H Commelinaceae
Lian. Thero.
Sol Cop 1
176 Commelina commuris L.
Trai thng
94
177 Cyanotis papilionacea Roem. & Schult.f. 178 Cyanotis vaga (Lour.) Roem. & Schult.f. 179 Murdannia giganteum ( Vahl.) Bruckner 180 Murdannia simplex (Vahl.) Brenan
H Convallariaceae
Bch trai Burman Bch trai hoang Trai cao Trai l hp Song bo Bm dy Kit Lc ta C cm Mao th st Mao th mnh Mao th trn di Mao th vy Bch u Cng Kerr Ha hong H bin ru
Thero. Thero. Thero. Thero. Chamae. Thero. Chamae. Thero. Chamae. Chamae. Thero. Thero. Chamae. Chamae. Chamae. Epi. Thero.
Cop 1 Cop 1 Sp Sp Sp Sol Cop 1 Sol Sp Sol Sol Cop 1 Sol Sp Cop 2 Sol Sol
181 Disporum cantonense ( Lour.) Merr.

H Cyperaceae
182 Bulbostylis densa (Wall.) Hand. Mazz. 183 Carex lindleyana Nees 184 Cyperus digitatus Roxb. 185 Cyperus halpan L. 186 Fimbristylis ferruginea (L.) Vahl. 187 Fimbristylis gracilenta Hance 188 Fimbristylis oblonga T.Koyama 189 Fimbristylis pubisquama Kern 190 Kyllinga melanosperma Nees 191 Scleria kerrii Turrill
H Orchidaceae
192 Dendrobium christyanum Reichb.f. 193 Habenaria ciliolaris (L.) Kraenzel

H Phormiaceae
194 Dianella ensifolia (L.) DC.

H Poaceae
Hng bi Trc tho lng C thm C l gng Mao cc hoa nh C may vng Xoang trc ng c Xoang trc sc Song on Tc hnh ht C chn nhn Tc hnh tm
95
Chamae. Chamae. Cryp. Cryp. Chamae. Chamae. Chamae. Chamae. Chamae. Thero. Thero. Chamae.
Cop 3 Cop 3 Sp Sp Sp Sp Cop 1 Cop 3 Sp Sp Cop 1 Sp
195 Arudinella setosa Trin. 196 Axonopus affinis A.Chase 197 Axonopus combressus (Sw.) Beauv. 198 Capillipedium parviflorum (R.Dr.) Stapf 199 Chrysopogon fulvus (Spreng.) Chiov. 200 Coelorachis pratensis (Bal.) A.Cam. 201 Coelorachis striata (Steud.) A. Cam. 202 Diectomis fastigiata (Sw.) Kunth 203 Digitaria abludens (Roem. & Sch.) Veldk 204 Digitaria radicosa (Prese) Miq. 205 Digitaria violascens Link
206 Eragrostis ferruginea (Thumb.) P.Beauv. 207 Eragrostis nigra Nees ex Steud 208 Eragrostis nutans (Retz.) Nees ex Steud 209 Eragrostis zeylanica Nees & Mey. 210 Erianthus arundinaceus (Retz.) Jeswiet. ex Heyne 211 Erianthus fastigiatus Nees ex Steud 212 Eulalia fimbriata (Hack.) Kuntze 213 Eulalia phaeothrix (Hack.) Kuntze 214 Eulalia speciosa (Deb.) Kuntze 215 Eulalia velutina (Hack.) Kuntze 216 Exotheca abyssinica (A. Rich.) A nders. 217 Garnotia barbulata (nees) Merr. 218 Ichnanthus vicinus (F.M.Bail.) Merr. 219 Imperata cylindrica (L.) P.Beauv. 220 Ischaemum barbatum Retz. 221 Miscanthus floridulus (Labill.) Warb. ex Schum & Laut. 222 Panicum hayatae A.Cam 223 Panicum notatum Retz. 224 Panicum repens L. 225 Paspalum orbiculare Forst. 226 Paspalum urvillei Steud. 227 Phragmites australis (Cav.) Trin. ex steud. 228 Rhynchelytrum repens (Willd.) C.E. Hubb. 229 Saccharum spontaneum L. 230 Saceiolepis indica (L.) Chase 231 Seraria parviflora (Poir.) Roem. & Schult. 232 Setaria pumila (Poir.) Roem. & Schult. 233 Sporobolus tenellus Bal. 234 Themeda arguens (L.) HacK 235 Themeda arundinacea (Roxb.) HacK. 236 Themeda caudata (Nees.) HacK. 237 Themeda triandra Forssk.
Tinh tho st Tinh tho en Tinh tho nghin Tinh tho tch lan Lau Mao phng chm Ct v ra Ct v lng vng Ct v p Ct v lng Ngoi gip Gt nm L tho gn C tranh Mm ru L sng K Hayata K ni C ng San trn Sao urville Sy nam Hng nhung C bng lau Bc nh ui chn ui chn nh X t mnh L nhn L sy C phao L tam hng
Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Chamae. Cryp. Chamae. Chamae. Chamae. Thero. Cryp. Chamae. Chamae. Chamae. Thero. Chamae. Thero. Chamae. Chamae. Thero. Thero. Chamae. Chamae. Chamae.
Sp Sp Sp Sp Sp Sp Sp Sp Sp Sp Sp Sol Sol Cop 3 Sol Sol Sp Cop 1 Sol Cop 1 Sol Sol Cop 2 Sp Sol Sp Sp Sol Sp Sol Cop 1 Cop 1
96
238 Thysanolaena maxima (Roxb.) Kuntze 239 Urochloa paspaloides J & C, Presl
H Smilacaceae
t, cht C ui nhn Kim cang lin hng Th phc linh Kim cang thon
Chamae. Thero. Lian. Lian. Lian.
Cop 1 Sol Cop 1 Cop 2 Cop 1
240 Smilax corbularia Kunth. 241 Smilax glabra wall.ex.Roxb. 242 Smilax lanceifolia Roxb.
H Zingiberaceae
243 Hedychium gardnerianum Rosc. 244 Zingiber rubens Roxb.
Ngi tin Gng
Cryp. Cryp.
Sol Cop 3
Trong : Mega. = Megaphanerophytes (cy g ln), Micro. = Microphanerophytes (cy g nh), Nano. = Nanophanerophytes (cy dng bi), Chamae. = Chamaephytes (cy lu nm), Thero. = Therophytes (cy mt nm), Lian. = Lianophanerophytes (cy leo), Cryp. = Cryptophytes (cy chi n), Epi. = Epiphytes (cy b sinh). Nh vy a dng loi kiu rng thng ba l (Pinus kesiya) mc t nhin Lm ng v vng ln cn c ghi nhn l 244 loi, thuc 68 h thc vt, 179 chi ca 4 ngnh: Lycopodiophyta, Polypodiophyta, Pinophyta v Magnoliophyta. C 3 h mc a dng cao: H Poaceae c 27 chi, 45 loi H Asteraceae c 22 chi, 25 loi H Fabaceae c 16 chi, 27 loi Trong khi c ti 30 h ch c 1 chi, 1 loi. Chi a dng loi nht l chi Desmodium (thuc h Fabaceae) c 8 loi, k l cc chi Eragrostis, Themeda ( thuc h Poaceae) v Quercus ( thuc h Fagaceae) u c 4 loi. V tnh trng loi: c 8 loi c ghi nhn trong sch Vit Nam, trong 4 loi thang VU v 4 loi thang EN, chim t l 3,28% s loi. a dng v dng sng c ghi nhn 8 dng sng vi t l: Megaphanerophytes (0,82%), Microphanerophytes (9,01%), Nanophanerophytes (18,44%), Chamaephytes (27,46%), Therophytes (27,05%), Lianophanerophytes (6,15%), Cryptophytes (6,56%), Epiphytes (4,51%). T kt qu ta c c biu sau:
BIU T L SNG CA RNG THNG BA L (PINUS KESIRA) MC T NHIN LM NG V VNG LN CN
T l %
97
30 25 20 15 10 5 0
.Mega. Micro. Nano. Chamae Thero. Lian. Cryp. Epi.
Dng sng
Tuy dng Megaphanerophyte ch chim 0,82%, song loi thng ba l l loi g ln l loi u th sinh thi, chng to nn kiu rng tha cy l kim rt c trng cho c cao nguyn Lm Vin v cao nguyn Di Linh Lm ng vi cu trc 3 tng n nh thnh phn loi kh phong ph, che ph t 40% - 50% v lng nh sng rt di do cho tng c bi, nn tng ny gm nhiu dng sng v chim u th v t l.
98
PHN IV. KT LUN V KIN NGH I. KT LUN

S dng phng php xp chng nghin cu rng thng ba l (Pinus kesiya) 3 a im trn 3 ai cao khc nhau xc nh kch thc thch hp tiu chun nghin cu rng thng ba l mc t nhin Lm ng v vng ln cn. Sau tin hnh nghin cu 20 tiu chun cc ai cao khc nhau t 800m n 2000m, kt qu nh sau: - Kch thc thch hp tiu chun nghin cu kiu rng thng ba l mc t nhin Lm ng v vng ln cn l: 35m x 35m. - Ch s a dng sinh hc thc vt chung cho kiu rng thng ba l Lm ng v vng ln cn: DMarg. = 3,76. - Thnh phn loi thc vt c mch kiu rng thng ba l mc t nhin gm 244 loi thuc 179 chi, 68 h ca 4 ngnh: Lycopodiophyta, Polypodiophyta, Pinophyta v Magnoliophyta. - nhiu c th loi cp SOC: 1 loi. Cp Cop3: 19 loi. Cp Cop2 loi. Cp Cop1. loi. Cp Sp: loi. Cp Sol: 58 loi. - C 8 loi c ghi trong sch Vit Nam (4 loi thang VU,4 loi thang EN), chim 3,28% tng s loi. - C 8 dng sng: Mega. (0,82%), Micro. (9,01%), Nano. (18,44%), Chamae. (27,46%), Thero. (27,05%), Lian. (6,15%), Cryp. (6,56%), Epi. (4,51%).
II. KIN NGH

i vi rng thng ba l trng ch c 2 tng: Tng g ln v tng c, loi rng ny tri qua qu trnh din th v dn dn hnh thnh tng g nh. Trong kinh doanh lm nghip nn to iu kin cho tng g nh pht trin, khng nn pht b cc loi g nh ny khi chng cn tui c kch thc cy bi nh, duy tr cu trc 3 tng bn vng v mt sinh thi hc, to iu kin cho n nh v thnh phn loi v vic bo v mi trng.
99
A SUGGESTION OF USING THE APPROPRIATE SIZE FOR PERQUADRATE AND ITS APPLICATION TO THE RESEARCH OF PLANT BIODIVERSITY OF THE NATURAL THREE LEAVED PINE FOREST DISTRIBUTED AMONG LAM DONG AND SUB AREAS.
Nguyen Duy Chinh Faculty of Biology, University of DaLat Huynh Kim Anh Faculty of Natural Science, University of Phu Yen
SUMMANY
The paper present a method of using superposed plots to identify the most suitable area of perquadrates for the research into the biodiversity of the natural three leaved Pine forests distributed among Lam Dong and its subareas. The sizes of the superposed plots are: 10m x 10m, 15m x 15m, 20m x 20m, 25m x 25m, 30m x 30m, 35m x 35m, 40m x 40m. The 35m x 35m sized perquadrates are suggeted to be the most appropriate and most pratical for researchs of plant biodiversity, especially thoes of the three leaved pine forests (Pinus kesiya) naturally grown in Lam Dong and the its subareas with the altitude from 800m to 2000m. In our researches, the total number of perquadrates is 20. Each perquadrate has a record of its own indices including: geological co-ordinate (latitude and longitude), alttitude, gradient, klinogeotropicsm, species composition, life form, number of individuals, Margalef index. From the results we have specific index of biodiversity for natural three-leaved pine forests, species composition and diversity of the life form in the forests. The everage Margalef index (DMarg) is 3,76.The species composition is rather rich and diverse, including 244 species, 179 genera,68 families of 4 vascular plant phyta (Lycopodiophyta, Polypodiophyta, Pinophyta, Magnoliophyta). There are 8 species which have been recorded in Red Data Book of Viet Nam. There are 8 life forms, including: Megaphanerophytes (0,82%), Microphanerophytes (9,01%), Nanophanerophytes (18,44%), Chamaephytes (27,46%), Therophytes (27,05%), Lianophanerophytes (6,15%). Cryptophytes (6,56), Epiphytes (4,51%).
KEY WORDS: Superposed plot, Perquadrate, Biodiversity, Margalef Index, Three leaved
pine forest, Species composition, Life form, Lam Dong.
100
TI LIU THAM KHO
1. Nguyn Hong Ngha, 1977. Bo tn ti nguyn di truyn thc vt rng. Nxb Nng Nghip, H Ni. 2. Nguyn Ngha Thn, 2007. Cc phng php nghin cu thc vt. Nxb i hc Quc gia H Ni, H Ni. 3. Nguyn Ngha Thn, 1997. Cm nang nghin cu a dng sinh vt. Nxb Nng Nghip, H Ni. 4. Nguyn Tin Bn, 1997. Cm nang tra cu v nhn bit cc h thc vt ht kn Vit
Nam. Nxb Nng Nghip, H Ni.
5. Nguyn Tin Bn (ch bin), 2003 v 2005. Danh lc cc loi thc vt Vit Nam (Tp II, Tp III). Nxb Nng Nghip, H Ni. 6. Nguyn Tin Bn (Ch tch Hi ng bin son), 2007. Sch Vit Nam (Phn II Thc vt). Nxb KHTN v CN, H Ni. 7. Phm Hong H, 1999. Cy c Vit Nam (Quyn I,II, III). Nxb Tr, Thnh ph H Ch Minh. 8. Thi Vn Trng, 1978. Thm thc vt rng Vit Nam. Nxb KH v KT, H Ni. 9. Trung tm nghin cu ti nguyn v mi trng. i hc Quc gia H Ni, 2001. Danh lc
cc loi thc vt Vit Nam. Nxb Nng Nghip, H Ni.
101

DDSH Thuc Vat Rung Thong 3 La Lam Dong

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DDSH Thuc Vat Rung Thong 3 La Lam Dong

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B GIO DC V O TO TRNG I HC LT

M TI B 2009 14 30 CH NHIM TI: ThS. NGUYN DUY CHNH

PHN III: KT QU NGHIN CU I. XC NH KCH THC THCH HP CA TIU CHUN

Kch thc xp chng

theo n kch thc 35m x 35m th gn nh khng tng , v trn th c mt im un

Selaginella monospora Spring

Quyn b n bo t Rng can x gn Rng bit x ui phng l si

Thero. Epi. Nano. Epi.

Asplenium affine Sw.

Brainea insignis ( HooK.) J. Smith

Crypsinus rhynchoplyllus (HooK.) Copel.

Pyrrosia lingua (Thunb.) Farw. Var. lingua

Rng tai chut li dao Epi. Seo g, chn x Cryp.

Pinus kesiya Royle ex Gordon

Lepidagathis hyalina Ness

Thero. Chamae. Chamae. Thero. Chamae. Chamae. Thero. Nano. Micro.

Centella asiatica (L.)Urb.

Mahonia nepalensis DC.

Virburnum coriaceum Bl.

Chloranthus japonicus Sieb.

Si nht Cp mc hnh sao

Craibiodendron stellatum (Pierre ex Lanees) W.W. Sm

Vaccinium iteophyllum Hance

Aporusa serrate Gagnep.

Tai ngh rng Trng qu ngh thut Chm cua

Micro. Nano. Nano.

Desmodium concinnum DC. Indigofera stachyodes Lindl.

Lithocarpur dealbatus (HooK.f. & Thoms.) Rehd. Quercus helferiana A.DC.

D trng Si Helfer. Long m Langbian Cha d Dum Trung b

Micro. Micro. Thero. Nano. Nano.

Gentiana langbianensis A, Chev. ex H. Smith

Elsholtzia blanda (Benth.) Benth.

Rubus annamensis Card.

Asparagus filicinus Buch Ham ex D.Don.

Lian. Thero. Thero. Chamae. Cryp. Chamae. Chamae.

Cyanotis papilionacea Roem. & Schult.f.

Fimbristylis oblonga T.Koyama

Chamae. Chamae. Chamae. Lian. Cryp. Cryp.

Smilax lanceifolia Roxb.

Hedychium gardnerianum Rosc. Zingiber rubens Roxb.

Adiantum stenochlamys Bak.

Rng nguyt s Rng bch hoa nht Rng vi ln

Cryp. Chamae Cryp.

Woodwardia Japonica (L.f.) J.Sm.

Microlepia platyphylla (D.Don) J.Smith

Paragramma banaensis (C. Chr). Ching

Pinus kesiya Royle ex Gordon

Pimpenella diversifolia DC.

Hoya macrophylla Blume.

H da l to C ct heo Bch nhung sa Miu nh Sn cc ni B ch, bp l ng sm Java Vt dai Si nht

Buddleia asiatica Lour.

Codonopsis Javanica (Bl.) HooK.f.

Virburnum coriaceum Bl.

Chloranthus japonicus Sieb.

Ca di xoan Nem l liu Sc daltonii B ngt hai mu Trng qu ngh thut Tp m l nh

Glochidion daltonii (Muel-Arg.) Kurz. Sauropus bicolor Craib

Desmodium concinnum DC. Flemingia lincata var. glutinosa Prain

D trng Si lng Mua lng

Micro. Micro. Nano.

Melastoma saigonense (Kuntze) Merr.

Hypoxis aurea Lour.

Cyanotis vaga (Lour.) Roem. & Schult.f.

Disporum cantonense ( Lour.) Merr.

Fimbristylis oblonga T.Koyama